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ButtERFliEs (LEpiDoptERƒ) oF Chizƒmi, PhEk DistRict, NƒgƒlƒÄD

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DÊÄʦك֫
:ŽƵƌŶĂůŽĨdŚƌĞĂƚĞŶĞĚdĂdžĂͮǁǁǁ͘ƚŚƌĞĂƚĞŶĞĚƚĂdžĂ͘ŽƌŐͮϭϮĞĐĞŵďĞƌϮϬϭϰͮϲ;ϭϯͿ͗ϲϱϵϯʹϲϲϯϰ
çãã›Ù¥½®›Ý;>›Ö®—ÊÖã›ÙƒͿÊ¥«®þƒÃ®͕W«›»®ÝãÙ®‘ã͕
Eƒ¦ƒ½ƒÄ—͕/ė®ƒ
ISSN 0974-7907 (Online)
/^^EϬϵϳϰͲϳϴϵϯ;WƌŝŶƚͿ
dƐŚĞƚƐŚŽůŽEĂƌŽϭΘ^ĂŶũĂLJ^ŽŶĚŚŝϮ
North East Network, P.O. Chizami, Phek District, Nagaland 797102, India
dŝƚůŝdƌƵƐƚ͕ϰϵZĂũƉƵƌZŽĂĚŶĐůĂǀĞ͕ŚŽƌĂŶ<ŚĂƐ͕ĞŚƌĂĚƵŶ͕hƩĂƌĂŬŚĂŶĚϮϰϴϬϬϭ͕/ŶĚŝĂ
1
tshetsholo@gmail.com, 2 sanjay.sondhi1@gmail.com (corresponding author)
1
2
KWE^^
ďƐƚƌĂĐƚ͗dŚĞďƵƩĞƌŇŝĞƐŽĨŚŝnjĂŵŝ͕WŚĞŬŝƐƚƌŝĐƚŽĨEĂŐĂůĂŶĚ͕/ŶĚŝĂǁĞƌĞĚŽĐƵŵĞŶƚĞĚŽǀĞƌĂƉĞƌŝŽĚŽĨƚŚƌĞĞLJĞĂƌƐ;ϮϬϭϭʹϮϬϭϰͿ͘dŚŝƐ
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;ϮϭϮͿƐƉĞĐŝĞƐǁĞƌĞƌĞĐŽƌĚĞĚŽĨǁŚŝĐŚϯϰƐƉĞĐŝĞƐĂƌĞůĞŐĂůůLJƉƌŽƚĞĐƚĞĚƵŶĚĞƌǀĂƌŝŽƵƐƐĐŚĞĚƵůĞƐŽĨƚŚĞtŝůĚůŝĨĞ;WƌŽƚĞĐƟŽŶͿĐƚ͕ϭϵϳϮ͘
ŵŽŶŐƐƚƚŚĞŵĂŶLJƐŝŐŶŝĮĐĂŶƚƌĞĐŽƌĚƐĚƵƌŝŶŐƚŚĞƐƵƌǀĞLJ͕ƚǁŽƐƉĞĐŝĞƐ͕ƚŚĞdĂǁŶLJŵƉĞƌŽƌChitoria ulupi ulupiĂŶĚƚŚĞŽŵŝĐKĂŬďůƵĞ
Arhopala comicaǁĞƌĞƌĞĐŽƌĚĞĚĨƌŽŵ/ŶĚŝĂĂŌĞƌĂƉĞƌŝŽĚŽĨϭϬϬLJĞĂƌƐ͘
<ĞLJǁŽƌĚƐ͗ƵƩĞƌŇLJĚŝǀĞƌƐŝƚLJ͕ĐŚĞĐŬůŝƐƚ͕ŚŝnjĂŵŝ͕Arhopala comica͕ŝŶǀĞŶƚŽƌLJ͕>ĞƉŝĚŽƉƚĞƌĂ͕EĂŐĂůĂŶĚ͕ƌĞĐŽƌĚ͕Chitoria ulupi.
Junonia almana almana
Peacock Pansy
NOT
EVALUATED
DATA
DEFICIENT
LEAST
CONCERN
NE
DD
LC
NEAR
VULNERABLE
THREATENED
NT
VU
ENDANGERED
CRITICALLY
ENDANGERED
EXTINCT
IN THE WILD
EXTINCT
EN
CR
EW
EX
K/͗ŚƩƉ͗ͬͬĚdž͘ĚŽŝ͘ŽƌŐͬϭϬ͘ϭϭϲϬϵͬ:Ždd͘Žϯϵϵϱ͘ϲϱϵϯͲϲϯϰͮŽŽĂŶŬ͗ƵƌŶ͗ůƐŝĚ͗njŽŽďĂŶŬ͘ŽƌŐ͗ƉƵď͗ϵϮ&ϱϳϲͲϯͲϰϮϮͲϰϯͲϴϭϳϮϯ&ϬϬ
ĚŝƚŽƌ͗ŶŽŶLJŵŝƚLJƌĞƋƵĞƐƚĞĚ͘
ĂƚĞŽĨƉƵďůŝĐĂƟŽŶ͗ϭϮĞĐĞŵďĞƌϮϬϭϰ;ŽŶůŝŶĞΘƉƌŝŶƚͿ
DĂŶƵƐĐƌŝƉƚĚĞƚĂŝůƐ͗DƐηŽϯϵϵϱͮZĞĐĞŝǀĞĚϭϮƉƌŝůϮϬϭϰͮ&ŝŶĂůƌĞĐĞŝǀĞĚϭϲKĐƚŽďĞƌϮϬϭϰͮ&ŝŶĂůůLJĂĐĐĞƉƚĞĚϮϴEŽǀĞŵďĞƌϮϬϭϰ
ŝƚĂƟŽŶ͗EĂƌŽ͕d͘Θ^͘^ŽŶĚŚŝ(2014). ƵƩĞƌŇŝĞƐ;>ĞƉŝĚŽƉƚĞƌĂͿŽĨŚŝnjĂŵŝ͕WŚĞŬŝƐƚƌŝĐƚ͕EĂŐĂůĂŶĚ͕/ŶĚŝĂ. Journal of Threatened Taxaϲ;ϭϯͿ͗ϲϱϵϯʹϲϲϯϰ; ŚƩƉ͗ͬͬ
Ědž͘ĚŽŝ͘ŽƌŐͬϭϬ͘ϭϭϲϬϵͬ:Ždd͘Žϯϵϵϱ͘ϲϱϵϯͲϲϯϰ
ŽƉLJƌŝŐŚƚ͗ ©EĂƌŽΘ^ŽŶĚŚŝϮϬϭϰ͘ƌĞĂƟǀĞŽŵŵŽŶƐƩƌŝďƵƟŽŶϰ͘Ϭ/ŶƚĞƌŶĂƟŽŶĂů>ŝĐĞŶƐĞ͘:ŽddĂůůŽǁƐƵŶƌĞƐƚƌŝĐƚĞĚƵƐĞŽĨƚŚŝƐĂƌƟĐůĞŝŶĂŶLJŵĞĚŝƵŵ͕ƌĞƉƌŽĚƵĐƟŽŶĂŶĚĚŝƐƚƌŝďƵƟŽŶďLJƉƌŽǀŝĚŝŶŐĂĚĞƋƵĂƚĞĐƌĞĚŝƚƚŽƚŚĞĂƵƚŚŽƌƐĂŶĚƚŚĞƐŽƵƌĐĞŽĨƉƵďůŝĐĂƟŽŶ͘
&ƵŶĚŝŶŐ͗ dŚĞEŽƌƚŚĂƐƚEĞƚǁŽƌŬ;EEͿǁŝƚŚƚŚĞƐƵƉƉŽƌƚŽĨΘDŝƐĞƌĞŽƌ͕'ĞƌŵĂŶLJ;ϮϬϭϬͲDĂƌĐŚϮϬϭϯͿĂŶĚEĂŐĂůĂŶĚtŝůĚůŝĨĞĂŶĚŝŽĚŝǀĞƌƐŝƚLJŽŶƐĞƌǀĂƟŽŶ
dƌƵƐƚ;DĂLJϮϬϭϯͲĞĐĞŵďĞƌϮϬϭϰͿĨƵŶĚĞĚEE͛ƐŽŶƐĞƌǀĂƟŽŶĚƵĐĂƟŽŶWƌŽŐƌĂŵŵĞĂŶĚĞŶĂďůĞĚ^ĂŶũĂLJ^ŽŶĚŚŝ͛ƐǀŝƐŝƚƚŽŚŝnjĂŵŝĂŶĚdƐŚĞƚƐŚŽůŽEĂƌŽǁŝƚŚƚŚĞ
,'ƚŽĚŽƚŚĞƐƚƵĚLJ͘
ŽŵƉĞƟŶŐ/ŶƚĞƌĞƐƚ͗dŚĞĂƵƚŚŽƌƐĚĞĐůĂƌĞŶŽĐŽŵƉĞƟŶŐŝŶƚĞƌĞƐƚƐ͘
ƵƚŚŽƌŽŶƚƌŝďƵƟŽŶƐ͗dEƐƵƌǀĞLJĞĚďƵƩĞƌŇŝĞƐŝŶƚŚĞĂƌĞĂĨƌŽŵϮϬϭϭƚŽϮϬϭϰ͘dEĂŶĚŵĞŵďĞƌƐŽĨƚŚĞ,'ƚŽŽŬŵŽƐƚŽĨƚŚĞƉŚŽƚŽŐƌĂƉŚƐŽĨƚŚŝƐƉĂƉĞƌ͘^^ŵĂĚĞ
ĮǀĞǀŝƐŝƚƐƚŽŚŝnjĂŵŝĂŶĚƐƵƌǀĞLJĞĚďƵƩĞƌŇŝĞƐĚƵƌŝŶŐŚŝƐǀŝƐŝƚƐ͘dEŝĚĞŶƟĮĞĚŵĂŶLJŽĨƚŚĞƐƉĞĐŝĞƐ͕ĂŶĚ^^ĐŽŶĮƌŵĞĚƚŚĞŝƌŝĚĞŶƟƟĞƐ͘^^ĐŽŶĚƵĐƚĞĚƚŚĞůŝƚĞƌĂƚƵƌĞ
ƌĞǀŝĞǁ͘dŚĞƉĂƉĞƌǁĂƐǁƌŝƩĞŶũŽŝŶƚůLJǁŝƚŚďŽƚŚĂƵƚŚŽƌƐĐŽŶƚƌŝďƵƟŶŐƚŚĞŝƌƌĞĐŽƌĚƐ͘dEƉƌĞƉĂƌĞĚƚŚĞƉůĂƚĞƐ͘
ƵƚŚŽƌĞƚĂŝůƐ͗ dÝ«›ãݫʽÊEƒÙÊ;ůŽͿŝƐĂLJŽƵŶŐŶĂƚƵƌĞĐŽŶƐĞƌǀĂƟŽŶŝƐƚďĂƐĞĚŝŶŚŝnjĂŵŝ͕WŚĞŬŝƐƚƌŝĐƚ͕EĂŐĂůĂŶĚĂŶĚĐƵƌƌĞŶƚůLJǁŽƌŬŝŶŐǁŝƚŚĂŶE'K͕EŽƌƚŚĂƐƚ
EĞƚǁŽƌŬ;EEͿ͕ŝŶŝƚƐEĂƚƵƌĞŽŶƐĞƌǀĂƟŽŶĚƵĐĂƟŽŶWƌŽŐƌĂŵŵĞǁŚŝĐŚŝŶǀŽůǀĞƐĐŽŶƐƚĂŶƚĞŶŐĂŐĞŵĞŶƚǁŝƚŚLJŽƵƚŚƐĂŶĚĐŚŝůĚƌĞŶƚŽǁĂƌĚƐĐƌĞĂƟŶŐĂĐŽŶƐĐŝŽƵƐŶĞƐƐ
ĨŽƌĐŽŶƐĞƌǀĂƟŽŶ͘ůŽŚĂƐĂŶĞŶƚŚƵƐŝĂƐƟĐŝŶƚĞƌĞƐƚŝŶƐƚƵĚLJŝŶŐďƵƩĞƌŇŝĞƐĂŶĚŚĂƐďĞĞŶŝĚĞŶƟĨLJŝŶŐďƵƩĞƌŇŝĞƐŝŶĂŶĚĂƌŽƵŶĚŚŝnjĂŵŝƐŝŶĐĞϮϬϭϭ͖ŚĞĂůƐŽŚĂƐŬĞĞŶ
interest in nature photography and videography. ^ƒÄ¹ƒù^Êė«®͕&ŽƵŶĚĞƌdƌƵƐƚĞĞdŝƚůŝdƌƵƐƚ͕ŝƐĂŶĂƚƵƌĂůŝƐƚǁŝƚŚĂŶŝŶƚĞƌĞƐƚŝŶ>ĞƉŝĚŽƉƚĞƌĂ͕ŚĞƌƉĞƚŽĨĂƵŶĂĂŶĚ
ĂǀŝĨĂƵŶĂ͘,ĞƐƵƉƉŽƌƚƐĐŽŶƐĞƌǀĂƟŽŶƌĞƐĞĂƌĐŚĂŶĚĂĐƟŽŶĂŶĚĐŽŶƐĞƌǀĂƟŽŶĞĚƵĐĂƟŽŶƉƌŽŐƌĂŵƐŝŶĞĂƐƚĞƌŶĂŶĚǁĞƐƚĞƌŶ,ŝŵĂůĂLJĂ͘
ĐŬŶŽǁůĞĚŐĞŵĞŶƚƐ͗dEƚŚĂŶŬƐƚŚĞEŽƌƚŚĂƐƚEĞƚǁŽƌŬĨŽƌƚŚĞŽƉƉŽƌƚƵŶŝƚLJƚŚĞLJŚĂǀĞŐŝǀĞŶŚŝŵƚŽŐĞƚŝŶǀŽůǀĞĚŝŶŶĂƚƵƌĞĐŽŶƐĞƌǀĂƟŽŶĞĚƵĐĂƟŽŶƉƌŽŐƌĂŵŵĞ
ĂŶĚŝŶŵĂŶLJŽƚŚĞƌǁĂLJƐ͘^^ĂĐŬŶŽǁůĞĚŐĞƐƚŚĞƐƵƉƉŽƌƚŐŝǀĞŶƚŽŚŝŵďLJEŽƌƚŚĂƐƚEĞƚǁŽƌŬĚƵƌŝŶŐŚŝƐǀĂƌŝŽƵƐǀŝƐŝƚƐƚŽEĂŐĂůĂŶĚ͘dŚĞĂƵƚŚŽƌƐǁŽƵůĚůŝŬĞƚŽƚŚĂŶŬ
ƚŚĞƌĞǀŝĞǁĞƌƐĨŽƌƚŚĞŝƌĨĞĞĚďĂĐŬĂŶĚĐŽŵŵĞŶƚƐ͘
6593
Butterflies of Chizami
N
C
Naro & Sondhi
records in this paper are through photographs taken by
members of the HGEC.
N
The hill state of Nagaland has a geographical
area of 16,579km2 of which 30% is under forest with
88% of this being private forests. Nagaland falls in
the eastern Himalaya region of the Himalaya Global
Biodiversity Hotspot, one of 34 global biodiversity
hotspots. Nagaland is home to a variety of forest types
(altitudes vary from 184–3,841 m) varying from northern
tropical wet evergreen forest, northern tropical semievergreen forest, northern sub-tropical broad-leaved
wet hill forest (500–1,800 m), northern sub-tropical
pine forest (1,000–1,500 m), northern montane wet
temperate forest (2,000–2,500 m) and temperate
forest (above 2,500m) (http://ibcn.in/wp-content/
uploads/2011/12/33-793_812- Nagaland.pdf).
Chizami is located in the Phek District in southeastern
Nagaland. Phek District, covering an area of 2,026km2
is bordered by Myanmar on the east and the state
of Manipur on the south. To its north are Kiphire and
Zunheboto districts, while Kohima District is to its west.
The village of Chizami is 88km due east of Kohima (Fig.
1).
It covers an area of 85,242ha approximately and the
altitudinal range varies from 981–2000 m. The forest
type at Chizami is mainly sub-tropical pine forest and
some northern sub-tropical broad-leaved wet hill forest.
The Chakhesang tribe inhabits Chizami. Chizami village
shares its boundary with Pholami villages on the north;
Khomi, Sumi and Losami Village on the northeast; Lai
Village of Manipur State on the east, Thetsumi Village on
the south; Enhulumi Village on the west and Porba and
Sakraba villages on the northwest. The Kuwari River is
to the east and the Tuphale River to the west of Chizami.
North East Network (NEN) (www.northeastnetwork.
org) is a women’s rights organization working in the
northeastern region of India with a focus on women’s
rights. NEN has also been working on the issue of natural
resource management in the area. As part of the Natural
Resource Management activity, wildlife and biodiversity
conservation education programme was launched in
2010 and the Hoolock Gibbon Eco Club was formed.
The Hoolock Gibbon Eco Club (HGEC) is a children’s
eco club consisting of 40 members from the villages of
Chizami and Enhulumi between the age group of 10–14
years. The HGEC is focused on conservation education,
conservation research and conservation action. As part
of the HGEC’s activities, primarily Tshetsholo Naro (TN),
Kewekhrozo Thopi, NEN and Sanjay Sondhi (SS), Titli
Trust (www.titlitrust.com), introduced the children to
butterfly watching and identification. Quite a few of the
S
The authors conducted butterfly surveys in the
area over a period of three years. The butterfly study
was mainly conducted in the following areas (Fig. 2).
(i) NEN compound (25024’0’’N & 94024’E, 1440m); (ii)
In and around Chizami Village; (iii) The road leading to
Thetsumi Village approximately 5km away from Chizami
Village; (iv) The Ewelu path up to approximately 3km
from Chizami Village; (v) Erule River bank - a mini hydro
project area 10km away from Chizami Village. Areas
that were primarily visited are listed below along with
photographs of their habitats (Appendix 1).
More than 50% of the butterflies seen were in the NEN
compound. TN lives in Chizami, and opportunistically
recorded and photographed butterflies in the area
beginning 2010. SS made five visits to Chizami in July
2011, February 2012, October 2012, January 2013
i ure
a of Na a and
he
is ri
Chizami
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
i ure
a of Chizami and surroundin areas
N N om ound
n and around Chizami i a e
he sumi oad
and March 2013 during which he surveyed butterflies
along with TN. The butterflies were photographed and
identified using existing literature (Marshall 1882–90;
Bingham 1905–07; Swinhoe 1912–13; Evans 1932,
1949; Talbot 1939, 1947; Wynter-Blyth 1957; Eliot 1969;
Corbet et. al 1992; Haribal 1992; Smith 1993; Smith
1994; Kunte 2000; Kinyon 2004; Kehimkar 2008; Singh
2011; Sondhi et. al 2013; Sondhi & Kunte 2014). Cryptic
species were netted and their external features relevant
for identification were observed and photographed.
Wherever possible, the identification of the individuals
were done up to the subspecies level. In numerous
species, there are different common names in use.
Alternate common names, if in use, are mentioned in
brackets.
S
S N
SC SS
N
Two-hundred-and-twelve (212) species were
recorded during the survey. The family Nymphalidae
with 102 species representing 48% of the species seen
was the most abundant family followed by Lycaenidae
with 36 species (17%) and Hesperiidae with 30 species
(14%). Of the 212 species recorded in Chizami, 34
species are legally protected under various schedules of
the Wildlife (Protection) Act, 1972 (Table 1).
Surveys and collections of butterflies from the Naga
Hills are scanty and Varshney’s bibliography (Varshney
2008, 2009) was consulted along with SS’s perusal of
eu a h
ru e i er
older literature. Amongst the earliest publications on
butterflies from the Naga Hills was the publication by
Elwes on the collection of butterflies by Mr. W. Doherty
during two expeditions in 1889 and 1890 (Elwes 1891).
In 1889, Doherty spent the rainy season collecting in the
Naga Hills from June to September. Subsequent to Mr.
Doherty’s butterfly collection in the Naga Hills, Major
H.C. Tytler spent many years collecting and reporting
on the butterflies of the Naga Hills (Tytler 1911, 1912).
Tytler’s collection covered the lower, medium and higher
altitudes of the Naga Hills covering areas between 200
and 2,100 m. Later, Tytler, who had by then become a
Lieutenant Colonel, was posted in Imphal in Manipur and
further collected butterflies from Manipur and Naga hills
(Tytler 1914, 1915a,b). In this paper, we will be referring
to only these species recorded from the Naga Hills.
Amongst the significant records during the survey,
two species, the Tawny Emperor Chitoria ulupi ulupi and
the Comic Oakblue Arhopala comica have been recorded
from India after 100 years, having last been recorded
during surveys by Tytler (1915a,b). The Comic Oakblue
Arhopala comica has previously only been known from a
few specimens collected in Manipur, hence this record
extends its range marginally to Nagaland. There are
numerous others species and subspecies for which there
exist no or very few recent published records from India
such as Assam Darter Ochlodes siva siva, Purple Swift
Caltoris tulsi, Ruddy Pied Flat Pseudocoladenia fatua,
Khaki Silverline Spindasis rukmini, Hedge Cupid Bothrinia
chennelli, Blue Duke Euthalia durga splendens, and
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
6595
Butterflies of Chizami
a e
Naro & Sondhi
S e ies re orded from Chizami o ered in
S hedu e
aenidae
Arhopala comica
Spindasis rukmini
N m ha idae
Bassarona durga splendens
Chitoria ulupi ulupi
Dilipia morgiana
Mimathyma chevana chevana
Sephisa Chandra Chandra
S hedu e
aenidae
Anthene lycaenina lycambes
Arhopala ganesa watsoni
Bothrinia chennelli
Euchrysops cnejus cnejus
Flos areste
Heliophorus moorei tytleri
Jamides elpis pseudoelpis
Lampides boeticus
Poritia hewitsoni hewitsoni
s hedu es
N m ha idae
Aemona amathusia
Calinaga brahma brahma
Euthalia franciae franciae
Euthalia aconthea garuda
Faunis eumeus assama
Hestina persimilis persimilis
Parthenos sylvia gambrisius
Kallima knyvettii
Lethe sinorix sinorix
Libythea lepita lepita
Melanitis zitenius zitenius
Mycalesis mestra sadona
Symbrenthia niphanda niphanda
ieridae
Appias albina darada
S hedu e
es eriidae
Hyarotis adrastus praba
N m ha idae
Euploea mulciber mulciber
Euthalia lubentina lubentina
ieridae
Appias galba
Green Duke Euthalia sahadeva nadaka. These species
are discussed in more detail in the notes below.
The present work is significant because it reports
numerous species not recorded by Tytler and Elwes
during their extensive collections of butterflies in the
Naga Hills. Equally significant, there are more than a
hundred species that were collected by them that were
not recorded at Chizami, though the survey area and
elevations covered during their surveys were far more
extensive.
N
S N
B
S
C
ami
es eriidae Su fami Coe iadinae
Bro n
Badamia exclamationis a ri ius
Recorded from the NEN compound in June 2012 and
July 2013 at the same location. Rare. Reported to be not
uncommon at Kohima (Tytler 1912).
reen
e Bibasis vasutana oore
Recorded from the NEN compound in October
2011. Rare. Reported from the Naga Hills (Elwes 1892).
Reported to be common at Kohima (Tytler 1912).
ndian
in Choaspes benjaminii japonica
urra
Recorded from near the stream on the Thetsumi
roadside at Chizami in July and November 2011. It
6596
should be noted that while this subspecies is the one
that is most commonly recorded in northeastern India,
species of this genera are best separated by examining
male genitalia. Rare. Reported from the Naga Hills
(Elwes 1892). Several individuals collected at Kohima
(Tytler 1912).
Sa e
Hasora anura anura de Ni i e
Recorded at NEN compound in November 2011.
Very rare.
hi e anded
Hasora taminatus bhavara
ruhs orfer
Recorded from the NEN compound in the month of
July 2012. Rare.
ami
es eriidae Su fami
es eriinae
B ue s otted S ru
o er Aeromachus kali de
Ni i e
Recorded on the Chizami road leading to Enhulumi
Village in May 2012. Rare. Reported to be common in
the Naga Hills (Elwes 1892; Tytler 1912).
eined S ru o er Aeromachus stigmata obsoleta
oore
Recorded from the NEN compound and the Erule
River at Chizami during the post-monsoon season.
Uncommon. Reported from the Naga Hills (Elwes 1892;
Tytler 1912).
ores
o
er Astictopterus jama olivascens
oore
Recorded from the Erule River and the Thetsumi trail
at Chizami between March and July in the pre-monsoon
season. Common. Reported from the Naga Hills (Elwes
1892; Tytler 1912).
ur e S i Caltoris tulsi de Ni i e
Recorded from the NEN compound in late November
2011. Few recent published records from India. Rare.
S rai h S i Parnara f guttatus Bremer & re
Recorded in the flower garden at NEN compound in
August 2011 and October 2012. Members of this genus
are best separated by male genitalia; hence this has only
been provisionally identified. Common. Reported from
the Naga Hills (Elwes 1892). Reported to be not common
at Kohima (Tytler 1912).
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
Sma Branded S i Pelopidas mathias mathias
a ri ius
Recorded from the NEN compound in the flower
garden in August 2011. Uncommon.
ar e Branded S i
Pelopidas sinensis
a i e
Recorded from the NEN compound in the month
of February 2012. Reported from the Naga Hills (Elwes
1892). Reported to be common at Kohima (Tytler 1912).
e o S o S i Polytremis eltola eltola e i son
Recorded from all locations at Chizami. Mostly seen
during the pre- and post- monsoon seasons. Common.
Reported to be common at Kohima (Tytler 1912).
ar s
Potanthus s
Numerous records from the Chizami area in the preand post-monsoon seasons. Numerous species reported
from the Naga Hills (Elwes 1892).
ssam ar er Ochlodes siva siva oore
Two individuals sighted by TN. Seen 400m away from
NEN office (West) in May 2010 and at the NEN compound
in May 2014. The first individual was a female, lacking
the UPF brand. However, UPH discal spots were as
opaque as the opaque spots in space 1b UPF and not
hyaline as are the other spots UPF, separating it from
Ochlodes brahma (Evans 1949). In addition, the spot in
1b in O. brahma is much smaller. Rare. These are the
first records of this species from Nagaland, having been
recorded from Khasi Hills in Meghalaya, Mussoorie in
Uttarakhand, Assam and north Myanmar (Evans 1949).
These sightings are also the first published record of this
species from India in many decades.
i h S ra
e Pithauria stramineipennis
stramineipennis ood ason & de Ni i e
Single record from Erule River bank at Chizami in
August 2013. SS had observed this species in the foothills
from other parts of northeastern India, while it has been
recorded up to 640m in Nepal (Smith 1994), hence this
record from 700m is unusual. Very rare in the survey
area. Reported from the Naga Hills at low elevations in
March and October (Tytler 1912).
u ed e Sebastonyma dolopia e i son
Recorded from the NEN compound in October 2012.
Rare in the survey area. Reported from the Naga Hills
(Tytler 1912).
ndian a m Bo Suastus gremius gremius a ri ius
Single record from Erule River bank at Chizami in
August 2013. Very rare in the survey area.
Si im a m ed e e Erionota torus ans
Recorded from all locations at Chizami. Separated
from similar-looking E. thrax by the rounded FW apex
and convex termen, though it is pertinent to note that
this species is best separated from E. thrax by male
genitalia. Mostly seen during the pre-monsoon season.
An individual entered the training room at night on
two consecutive nights in July 2011, evidently attracted
to light. Common. Erionota thrax reported from the
Naga Hills with individuals captured at Kohima inside a
bungalow, apparently attracted to light, just as we had
observed (Tytler 1912). However, Evans described E.
torus as a separate species from E. thrax only in 1941,
hence Tytler’s records could have been either of these
two species.
Ba
eined Branded ed e e Matapa cf. sasivarna
oore
Recorded on the Chizami road leading to Thetsumi
in October. The upperside of this individual was
not observed. This species could be M. cresta, M.
purpurascens or M. sasivarna. In the field, these species
are best separated by a combination of the male UPF
brands and wing markings and coloration. While in M.
cresta and M. purpurascens the male brand is black and
inconspicuous, in M. sasivarna, it is grey and conspicuous.
However, on the basis of the absence of paler UNF apex
and UNH base, present in M. cresta, and the metallicgreen scales at HW base and wing coloration, this is
provisionally identified as M. sasivarna (de Jong 1983;
Evans 1949). Very rare.
es ri ed emon Notocrypta curvifascia curvifascia
e der & e der
Recorded from all locations at Chizami. Mostly seen
in the post-monsoon season. Common. Reported to be
common at Kohima (Tytler 1911).
S otted emon Notocrypta feisthamelii alysos
oore
Recorded on the Chizami to Awelu trail near the
stream in the pre-monsoon season. Uncommon.
Reported to be common at Kohima (Tytler 1912).
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
Common Banded emon Notocrypta paralysos
asawa ruhs rofer
Single record from the NEN compound in July 2011.
Rare in the survey area.
ree itter Hyarotis adrastus praba oore
Recorded on the Chizami to Erule trail in the premonsoon season. Uncommon. This subspecies is legally
protected under Schedule IV of the Wildlife (Protection)
Act, 1972. Reported from the Naga Hills (Tytler 1912).
ami
es eriidae Su fami
r inae
Common S otted a Celaenorrhinus leucocera
o ar
Worn individuals were recorded from the NEN
compound in October 2012. Uncommon. Reported
from the Naga Hills in September (Tytler 1912).
udd
ied
a Pseudocoladenia cf. fatua
a n
n e Ctenoptilum vasava vasava
oore
Recorded from the Erule River at Chizami in March
2012. Rare. A single male collected from Nichuguard in
the Naga Hills (Tytler 1912).
Ches nu n e Odontoptilum angulata angulata
e der
Recorded from the Erule River at Chizami in March
2013. Rare in the survey area. Reported to be common
from the Naga Hills at low elevations (Tytler 1912).
ami
aenidae Su fami Cure nae
Bri h Sun eam Curetis bulis bulis es ood
Recorded two individuals at the NEN compound and
on the Erule trail at Chizami in October 2012. Rare in the
survey area. Reported from the Naga Hills (Elwes 1892;
Tytler 1912).
ans
ami
Recorded from the Thetsumi roadside near the stream
and at the NEN compound in May and June. Rare. This
species most closely resembles P. fatua with the spot in
two extended to beyond origin of vein 3, UPF, and being
brighter, similar to the West Himalayan species P. fatih
and its wings being more produced than P. festa which
also occurs at similar altitudes (Evans 1949). The similar
looking P. dan is more of a low elevation species. It is
pertinent to note that while Evans (1949) had treated
fatua and festa as a subspecies of Pseudocoladenia dan,
subsequent work has elevated both these subspecies to
species level (Huang & Xue, 2004). However, P. fatua
and P. festa are best separated by differences in male
genitalia, hence this species is provisionally identified.
Reported as Coladenia faith from Kohima in the Naga
Hills in September and October between 1,800 and 2,130
m (Tytler 1915b), and possibly represents the species P.
fatua as described by Evans (1949).
S otted Sno
a Tagiades menaka menaka
oore
Recorded from the NEN compound in October 2011.
Rare. Reported from the Naga Hills (Elwes 1892; Tytler
1912).
aenidae Su fami
aeninae
o den Sa hire Heliophorus brahma brahma
oore
Recorded from all locations at Chizami, throughout
the year, though less common in the winter months.
Very common. Reported to be common in the Naga Hills
(Elwes 1892; Tytler 1912).
Na a Sa hire Heliophorus kohimensis kohimensis
er
Recorded from only two locations at Chizami. On
the Ezilu road to Enhulumi and around NEN compound.
Both males and females seen at NEN compound. Seen
from May to October. Uncommon. Described as a new
species by Tytler from the Naga Hills (1911).
reen Sa hire
zure Sa hire Heliophorus
moorei tytleri i e
Recorded from the Government High school area and
NEN compound. Very common. This species is legally
protected under Schedule II of the Wildlife (Protection)
Act, 1972. Reported as very abundant in the Naga Hills
(Elwes 1892). Tytler (1911) reports it to be the most
common of the genus in the Naga Hills.
ami
S otted n e Caprona agama agama oore
Recorded from the Erule River at Chizami in March
2012. Rare.
aenidae Su fami o omma nae
oin ed Ci ia e B ue Anthene lycaenina lycambes
e i son
A single record from the Erule River at Chizami in
July 2012. Seen mud-puddling at the riverside. Very
rare in the survey area. This species is legally protected
under Schedule II of the Wildlife (Protection) Act, 1972.
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Reported from the Naga Hills (Elwes 1892; Tytler 1912).
Common ed e B ue Acytolepis puspa gisca
ruhs orfer
Recorded from all locations at Chizami. Very
common. Reported to be very common from the Naga
Hills (Elwes 1892; Tytler 1912).
ain ed e B ue Celastrina lavendularis limbata
oore
A single record by SS in July 2011. Very rare in the
survey area. Reported from the Naga Hills (Elwes 1892).
a e ed e B ue Udara dilecta dilecta oore
Recorded from the NEN compound and Erule River
at Chizami. Only two individuals seen in October 2011.
Rare in the survey area. Reported from the Naga Hills
(Elwes 1892).
ed e Cu id Bothrinia chennelli de Ni i e
Recorded from the Erule trail at Chizami. Seen in
October 2011 (TN) and March 2013 (SS). Rare. This
species is legally protected under Schedule II of the
Wildlife (Protection) Act, 1972. Reported from the Naga
Hills (Elwes 1892). Reported to be not common by Tytler
(1911).
Cha man s Cu id Everes argiades diporides
Cha man
Single record from the NEN compound at Chizami in
the month of October 2011. Very rare in the survey area.
Reported from the Naga Hills (Elwes 1892; Tytler 1912).
ndian Cu id Everes lacturnus assamica
er
Recorded from the Erule River and the NEN
compound in the pre- and post-monsoon seasons. Rare
in the survey area. Reported from the Naga Hills (Tytler
1912).
Si er or e me no Catochrysops panormus
exiguus is an
Recorded from two locations at Chizami; the NEN
compound and on the Erule trail. Seen mostly during
summer. Rare in the survey area. Reported from the
Naga Hills (Tytler 1912).
or e me no Catochrysops strabo strabo a ri ius
Recorded from the NEN compound. Two individuals
seen. Rare in the survey area. Reported from the Naga
Hills (Elwes 1892; Tytler 1912).
Naro & Sondhi
ram B ue Euchrysops cnejus cnejus a ri ius
A single record from the NEN compound at Chizami
in May 2012. Very rare in the survey area. This species
is legally protected under Schedule II of the Wildlife
(Protection) Act, 1972.
ar Ceru ean Jamides bochus bochus S o
Recorded from all locations at Chizami. Mostly seen in
May. Very common. Reported from the Naga Hills (Elwes
1892; Tytler 1912).
Ceru ean Jamides s e ies in S
A butterfly was observed by TN at Chizami in
November 2012, which looked like a Jamides in DSF and
could not be identified to the species level. The upperside
markings were not observed in the field. This individual
looked superficially like J. celeno in DSF, but the UNH
discal markings were narrower and broken, somewhat
like a Nacaduba species, compared with individuals
observed by SS in other areas of the Himalayas.
is enin Ceru ean Jamides elpis pseudoelpis
Bu er
Recorded at Chizami near the village (east of
Chizami). A single record in the month of January 2011.
Very rare in the survey area. This species is legally
protected under Schedule II of the Wildlife (Protection)
Act, 1972. Reported to be common from the Naga Hills
(Elwes 1892; Tytler 1912).
ea B ue Lampides boeticus innaeus
Recorded from all locations at Chizami. Most
common in the pre-monsoon season. Visits flowers.
Very common. This species is legally protected under
Schedule II of the Wildlife (Protection) Act, 1972.
Reported to be very common from the Naga Hills (Tytler
1912).
e ra B ue Leptotes plinius plinius a ri ius
Recorded from all locations at Chizami in April and
May. Common. A single individual collected at Kohima
(Tytler 1912).
in
ine ue Petrelaea dana de Ni i e
A single record from the Erule riverside at Chizami in
October 2011. Very rare in the survey area. Reported
from the Naga Hills (Tytler 1912).
Common ine ue Prosotas nora ardates
oore
Mostly recorded from the Erule riverside, mud-
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
6599
Butterflies of Chizami
Naro & Sondhi
puddling in groups in the summer. Very common.
Reported sparingly from 600m altitude in the Naga Hills
(Tytler 1912).
in Manipur. This species is legally protected under
Schedule I of the Wildlife (Protection) Act, 1972. Rare.
ai ess Bush ue Arhopala ganesa watsoni
ar rass B ue Zizeeria karsandra oore
Recorded from the NEN compound. Mostly seen
during summer. Common. Reported from the Naga Hills
(Tytler 1912).
ami
aenidae Su fami ori inae
Common em Poritia hewitsoni hewitsoni
oore
Recorded at Government High School area, Tahulu
trail at Chizami. Recorded two individuals at different
locations in the month of June 2012. This subspecies
is legally protected under Schedule II of the Wildlife
(Protection) Act, 1972. Rare. A few individuals collected
from Kohima (Tytler 1912).
ami
aenidae Su fami he inae
eaf B ue Amblypodia anita dina ruhs rofer
This species was recorded on the Erule trail and mostly
seen mud-puddling near the riverside. Subspecies level
identity could not be confirmed; this could also be the
ssp. gigantea, known from Manipur. Rare.
Bi s o o a Ancema ctesia ctesia e i son
A single record from the NEN compound in
September 2013. Very rare in the survey area. Reported
to be common at 1,500m in the Naga Hills (Tytler 1912).
ans
A single record from the Thetsumi roadside at
Chizami in January 2013. Very rare in the survey area.
This species is legally protected under Schedule II of the
Wildlife (Protection) Act, 1972.
e o dis
a
ue Arhopala singla de Ni
i e
Single record from the New Town area at Chizami in
September 2013. Very rare in the survey area. Reported
from the Naga Hills (Elwes 1892; Tytler 1912).
ai ess ush ue Flos areste e i son
Recorded from the NEN compound and the Erule
trail at Chizami in November 2011 and January 2013.
Four individuals sighted. Rare. This species is legally
protected under Schedule II of the Wildlife (Protection)
Act, 1972.
Common inse Catapaecilma major major ru e
Recorded only once at the Erule riverbank at Chizami
in March 2013. Very rare in the survey area. Reported
from the Naga Hills (Tytler 1912).
Si ers rea B ue Iraota timoleon timoleon S o
o dered a ue Arhopala bazalus teetsa de
i e
A single record from NEN compound in November
2013. Very rare in the survey area. Reported from the
Naga Hills (Elwes 1892).
Recorded two individuals from the NEN compound
in May 2013. Very rare in the survey area. Reported
from the Naga Hills in May (Tytler 1915b).
Comi a ue Arhopala comica de Ni i e
Two records by TN from the NEN compound on 1
April 2013 and 12 April 2014. Very rare. This Arhopala
species is easily told by the markings on the underside
as well as the shape of the hindwing tornus and its tail
(de Nicéville, 1900). The type locality of this species is
Bhamo in Myanmar while it was recorded as Surendra
learmondi (=Arhopala comica) from Loimwe in the
Southern Shan States (Tytler, 1940). From India, two
females of this species were recorded from Sebong,
Manipur in March (Tytler 1915b; Evans 1957). Hence this
record of Arhopala comica marginally extends the range
of this species into Nagaland, and is the first record from
India in a hundred years, since Tytler’s (1915b) collection
Recorded from the NEN compound and the Erule
trail in the pre-monsoon season from February to July.
Rare. Reported from the Naga Hills (Elwes 1892) where
it is not as common as in the Khasi Hills. Tytler (1911)
reports it to be very common in Kohima throughout the
year.
Ni
Common
ash Rapala nissa nissoides S inhoe
ha i Si er ine Spindasis rukmini de Ni i e
Recorded from the North East Network office
compound. Single sighting in October 2012. Very rare.
This species is legally protected under Schedule I of
the Wildlife (Protection) Act, 1972. A single individual
collected from the Naga Hills (Tytler 1912).
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
C u Si er ine Spindasis syama peguanus
Naro & Sondhi
oore
Recorded from the Government High school area and
on the Erule trail at Chizami in the pre-monsoon season.
Rare in the survey area. Reported from the Naga Hills
(Elwes 1892; Tytler 1912).
Common
a ia B ue Surendra quercetorum
quercetorum oore
Recorded from many different locations at Chizami
through most of the year. Very common. Reported
from the Naga Hills (Elwes 1892; Tytler 1912).
ami N m ha idae Su fami
a urinae
ur e m eror Apatura ambica ambica
o ar
Recorded from the Thetsumi trail and Erule riverside
at Chizami. Seen sitting in groups, mud-puddling.
Common. Reported to be common in the Naga foothills
up to 1,370m (Tytler 1911).
a n m eror Chitoria ulupi ulupi oher
Recorded by TN from the NEN compound on 14
June 2013 at 12:57hr. Seen sitting on human clothes.
This species is a powerful flier. There have been no
published records of this species from India in more
than a hundred years, having been last reported from
India in 1915 (Tytler 1915a). Very rare. The subspecies
florenciae, which is now synonymized with subspecies
ulupi, is legally protected under Schedule I of the Wildlife
(Protection) Act, 1972. Reported to be rare in the Naga
Hills (Tytler 1911, 1915a). Tytler (1911) initially believed
this to be a separate species from C. ulupi but later
stated (Tytler 1915a) that it would turn out to be a race
of the same species.
o den m eror Dilipia morgiana es ood
Recorded from the Thetsumi trail and the Erule
riverside at Chizami in October 2012 and August 2013
respectively. Common. Seen visiting an animal shed.
This species is legally protected under Schedule I of the
Wildlife (Protection) Act, 1972. Reported to be rather
rare in the Naga hills (Tytler 1911).
Ser ean m eror Mimathyma chevana chevana
oore
Only two individuals recorded from the NEN
compound and Erule riverside at Chizami in October
2012. Rare. This subspecies is legally protected under
Schedule I of the Wildlife (Protection) Act, 1972. A single
individual recorded from the Naga Hills (Tytler 1911).
Cour esan Euripus nyctelius nyctelius
ou eda
Recorded from Khili and inside the village at Chizami in
September and October 2011 respectively. Uncommon.
Reported to be common in the Naga Hills (Tytler 1911).
asha Herona marathus marathus ou eda
A single record from Sitikho at Chizami in October
2011. Rare.
Cir e Hestinalis nama nama ou eda
Recorded from many different locations at Chizami
in the pre- and post-monsoon seasons. Very common.
Reported to be very common in the Naga Hills (Tytler
1911).
Siren Hestina persimilis persimilis es ood
Two individuals recorded from NEN compound and
Chizami village in Aug 2011 and July 2013 respectively.
This species is legally protected under Schedule II of the
Wildlife (Protection) Act, 1972. Rare. A single individual
recorded in October in the Naga Hills (Tytler 1911).
Ba
rin e Rohana parisatis parisatis
es
ood
Recorded from the Erule riverside at Chizami in
October 2012. Seen sitting on moist stones and mudpuddling. Common. Males reported to be exceedingly
common in the Naga Hills (Tytler 1911).
as ern Cour er Sephisa chandra chandra
oore
Recorded from two locations at Chizami-the
Thetsumi trail and the Erule riverside in October 2011
and October 2012. Rare. This subspecies is legally
protected under Schedule I of the Wildlife (Protection)
Act, 1972. Numerous records from the Naga Hills (Tytler
1911).
ami N m ha idae Su fami Bi idinae
n ed Cas or Ariadne ariadne pallidior ruhs orfer
Recorded by SS from Chizami in October 2012. Rare
in the survey area.
Common Cas or Ariadne merione tapestrina
oore
A single record from Etsoshulu at Chizami in October
2012. Very rare in the survey area. Only one individual
collected from the Naga Hills (Tytler 1911).
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
ami N m ha idae Su fami Chara inae
Common Na a Charaxes athamas athamas rur
Rare in the survey area. Reported to be common near
Kohima from August to October (Tytler 1911).
Recorded from the Erule riverside at Chizami.
Seen mud-puddling during the summer. Uncommon.
Reported to be common from the Naga Hills up to
1,500m (Tytler 1911).
Cho o a e i er Parantica melaneus plataniston
ruhs orfer
Recorded from many different locations at Chizami.
Common. Reported to be not common; collected near
Kohima from August to October (Tytler 1911).
rea Na a Charaxes eudamippus eudamippus
ou eda
Recorded two individuals from NEN compound and
Erule riverside at Chizami in August 2011 and August
2013. Rare. Reported to be not uncommon in the Naga
foothills (Tytler 1911).
ami N m ha idae Su fami C res nae
Common
a Cyrestis thyodamas thyodamas
Boisdu a
Recorded by SS in January 2013, but no photographic
evidence. Rare in the survey area. Only a single individual
collected, not common in the Naga Hills (Tytler 1911).
ami N m ha idae Su fami
anainae
ain i er Danaus chrysippus chrysippus innaeus
Records by SS in flight at Chizami in July 2011 and
October 2012. Outside of Chizami, recorded by TN
at Pungro, Kiphire District, Nagaland in the month of
October 2013 where he sighted 2–3 individuals. Rare
in the survey area. A few individuals reported from the
foothills of the Naga Hills (Tytler 1911).
S ri ed i er Common i er Danaus genutia
genutia Cramer
Recorded from all locations at Chizami. Mostly seen
during summer. Common. Reported from the Naga
Hills from the foothills up to 1,500m throughout the year
(Tytler 1911).
S ri ed B ue Cro Euploea mulciber mulciber
Cramer
Males and females recorded from the NEN
compound and the Thetsumi trail at Chizami in the preand post-monsoon seasons. Uncommon. This species is
legally protected under the Schedule IV of the Wildlife
(Protection) Act, 1972. Reported as very common in the
Naga Hills (Tytler 1911).
ass
i er Parantica aglea melanoides
oore
Single record on the road linking Kuwari and Chizami.
Ches nu i er Parantica sita sita o ar
Recorded from many different locations at Chizami.
Common. Reported to be common near Kohima from
August to October (Tytler 1911).
ar B ue i er Tirumala septentrionis septentrionis
Bu er
Recorded from all locations at Chizami between
April and October. Common. Reported to be common;
collected near Kohima from August to October and in
the foothills in February and March (Tytler 1911).
ami N m ha idae Su fami
e i oniinae
e o Cos er Acraea issoria issoria
ner
Recorded from many different locations at Chizami.
Mostly seen during the summer. Very common.
Reported to be very common from the Naga Hills in the
post-monsoon season (Tytler 1911).
ar e Si ers ri e Argynnis childreni childreni Gray
Recorded from all locations at Chizami between
March and October. Visits flowers. Very common.
Reported to be fairly common from the Naga Hills (Tytler
1911).
ro i a
ri ar
ndian ri ar
Argynnis
hyperbius hyperbius innaeus
Recorded from all locations at Chizami. Both males
and females (less common) seen between March and
October. Common. Reported to be very common from
the Naga Hills (Tytler 1911).
ed a e in Cethosia biblis tisamena ruhs orfer
Recorded from all locations at Chizami between
March and October. Very common. Reported to be
exceedingly common from the Naga Hills (Tytler 1911).
eo ard a e in Cethosia cyane cyane rur
Recorded from many different locations at Chizami.
Mostly seen visiting flowers during summer. Uncommon.
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
Reported to be not common from the Naga Hills (Tytler
1911).
ar e eoman Cirrochroa aoris aoris
ou eda
Recorded from many different locations at Chizami
between March and October. Seen visiting animal
droppings and flowers. Common. Reported to be not
very common from the Naga Hills (Tytler 1911).
Common eoman Cirrochroa tyche mithila
oore
Single record from the NEN compound in November
2011. Rare in the survey area. Reported to be not
uncommon from the Naga Hills (Tytler 1911).
us Cupha erymanthis lotis Su zer
Single record from Kuwari at Chizami in November
2013. Very rare in the survey area. Reported to be
rather scarce from the Naga Hills (Tytler 1911).
Common eo ard Phalanta phalantha phalantha
rur
Recorded from the NEN compound and the Erule
riverside at Chizami. Two records from these two
locations, seen during the pre-monsoon season. Rare in
the survey area. Reported to be common from the Naga
Hills (Tytler 1911).
a ran Vagrans egista sinha o ar
Recorded from the Erule riverside at Chizami in the
summer months. Seen visiting animal droppings and
mud-puddling. Rare in the survey area. Reported to be
common at low elevations from the Naga Hills (Tytler
1911).
Cruiser Vindula erota erota a ri ius
Recorded from Chizami. Both males and females
(less common) seen between March and October. Seen
visiting flowers. Uncommon. Males reported to be fairly
common from the Naga Hills (Tytler 1911).
ami N m ha idae Su fami i heinae
Common Bea Libythea lepita lepita oore
Two individuals recorded from the NEN compound
and the Erule River at Chizami in October 2012. Rare in
the survey area. This species is legally protected under
Schedule II of the Wildlife (Protection) Act, 1972. A few
individuals collected from the Naga Hills (Tytler 1911).
ami N m ha idae Su fami imeni dinae
ima a an Ser ean Athyma opalina orientalis
es
Several individuals recorded from two locations at
the Thetsumi roadside at Chizami between March and
October. Uncommon. Reported to be not uncommon
from the Naga Hills (Elwes 1891; Tytler 1911).
Common Ser ean Athyma perius perius innaeus
Recorded from all locations at Chizami between
March and October. Common. Reported to be very
common from the Naga Hills (Tytler 1911).
ran e S a Ser ean Athyma cama cama
oore
Recorded from the NEN compound in October 2012.
Both male and female seen. Rare. Reported to be not
uncommon from the Naga Hills (Tytler 1911).
S a Ser ean Athyma selenophora selenophora
o ar
Two records from the NEN compound and the Erule
trail at Chizami in October 2011. Rare. Reported to be
very common from the Naga Hills (Tytler 1911).
rea Calinaga brahma brahma Bu er
Recorded from the NEN compound. Two individuals
seen at the same location in May 2011 and March 2012.
Rare. This species is legally protected under Schedule II
of the Wildlife (Protection) Act, 1972. A few individuals
collected in April and May, near Kohima (Tytler 1911).
B ue u e Bassarona durga splendens
er
Recorded from Kuwari at Chizami in November
2013. Seen mud-puddling at the riverside. This is the
rare subspecies splendens in which the blue discal band
forms lunules throughout its length from tornus to costa
described by Tytler (1915a) from the Naga Hills. Very
rare. This species is legally protected under Schedule I of
the Wildlife (Protection) Act, 1972. Reported from near
Imphal in Manipur (Tytler 1915a).
ren h u e Euthalia franciae franciae e der &
e der
Mostly recorded from the Thetsumi trail at Chizami.
Common between July and October. This species is
legally protected under Schedule II of the Wildlife
(Protection) Act, 1972. Reported from the Naga Hills
(Elwes 1891). It is reported to be not uncommon in
August to October between 1,500–1,600 m in the Naga
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
Hills (Tytler 1911).
Bronze u e Euthalia nara nagaensis
er
A single record on the village footpath leading to
the NEN office in September 2012. Very rare. Reported
from the Naga Hills (Elwes 1891; Tytler 1911).
reen u e Euthalia sahadeva nadaka ruhs orfer
A male recorded basking high up on a tree in July
2011, below the NEN compound. Females recorded in
the summer in the NEN compound. Common. Reported
to be not uncommon from the Naga Hills (Elwes 1891;
Tytler 1911; Tytler 1915a).
Common Baron Euthalia aconthea garuda
oore
Recorded from many different locations at Chizamiboth males and females seen. Recorded in the pre- and
post-monsoon seasons. Common. This species is legally
protected under Schedule II of the Wildlife (Protection)
Act, 1972. Reported to be not common from the Naga
Hills (Tytler 1911).
aud Baron Euthalia lubentina lubentina Cramer
A single record from the Chizami Village in March
2012. Very rare in the survey area. This species is legally
protected under Schedule IV of the Wildlife (Protection)
Act, 1972. Only a few individuals collected in the Naga
Hills (Tytler 1911).
hi e ed ed B ue Baron Euthalia phemius
phemius ou eda
A single record of a male from the NEN compound in
October 2011. Very rare in the survey area. Reported
from the Naga Hills (Tytler 1911).
e o Sai er Neptis ananta ochracea ans
Two individuals recorded from the Thopfu trail and
the NEN compound in Sep 2013. Rare in the survey
area. Reported to be not uncommon in the Naga Hills
from July to October (Tytler 1911).
Nam a Sai er Neptis f namba namba
er
A single unconfirmed record from the NEN compound
in Oct 2011. Generally N. namba has brighter orange
markings compared with N. ananta (Eliot, 1969), but
this is not always true based on SS’s experience in the
field as well as images available online http://www.
ifoundbutterflies.org/2773-Neptis/Neptis-namba and
http://www.ifoundbutterflies.org/2770-Neptis/Neptisananta wherein some individuals of N. namba have paler
orange markings. While this individual was worn, and
its cilia not clearly visible, the UNF/H bluish tinge on the
markings below, so typical of namba was clearly visible
hence we are inclined to provisionally identify this as N.
namba. Neptis namba and Neptis ananta have often
been considered as a single species and forms/subspecies
of the latter. While Eliot suggested these were separate
species, he also stated “dissections of the male genitalia
have not helped greatly to establish whether there are
one, two or three species in the complex.” Clearly more
work on the phylogeny and ecology of these species are
necessary to establish their true status. Neptis namba
was described by Tytler (1915a) as a new species on the
basis of his collection from the Naga Hills. Tytler (1915a)
states that N. ananta is mostly found between 1,500–
2,100 m, while N. namba is mostly found in the foothills.
However, SS’s records from the NE India show that while
N. ananta is generally found at higher altitudes, both
species have been recorded from the foothills at least
up to 600m. However this record, if correct, extends its
altitudinal range even further.
Broad anded Sai er Neptis sankara amba
oore
Two individuals recorded from NEN compound in
June 2012 and August 2012. Rare.
a as s Sai er Neptis sappho astola oore
Recorded from all locations at Chizami between
March and October. Very common. Reported to be very
common from the Naga Hills (Tytler 1911).
Commander Moduza procris procris Cramer
A single record in flight by TN from the Erule River at
Chizami in 2012. Very rare in the survey area.
C i er Parthenos sylvia gambrisius a ri ius
A single record from the NEN compound in November
2011. Very rare in the survey area. This species is legally
protected under Schedule II of the Wildlife (Protection)
Act, 1972. Reported to be fairly common at low
elevations in the Naga Hills (Tytler 1911).
ami N m ha idae Su fami N m ha inae
ndian or oiseshe Aglais caschmirensis aesis
ruhs orfer
Recorded from many different locations at Chizami
in the pre- and post-monsoon seasons. Common. This
species has extended its range eastwards over time from
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
the western and central Himalaya and has established
a known population in western Arunachal Pradesh and
Nagaland (Naro 2012).
on o Araschnia prorsoides dohertyi B an hard
Recorded only from Thetsumi trail near the Tsoyiphe
stream in July 2011 and from Ezilu in March 2011 at
Chizami. Some authors treat Araschnia dohertyi as a
separate species. Rare. Reported to be abundant from
the Naga Hills between 1,800–2,400 m (Elwes 1891) who
states that its larval food plant is a nettle. Tytler (1911)
states that it is fairly common post-monsoon.
u umn eaf Doleschallia bisaltide indica
oore
Recorded from the NEN compound and the Erule trail
at Chizami, seen during summer season. Uncommon.
Common at low elevations but scarce near Kohima
(Tytler 1911).
ran e a eaf Kallima inachus inachus Boisdu a
Recorded from the Erule riverside at Chizami
between February and July. Common. Reported to be
fairly common from the Naga Hills (Elwes 1891; Tytler
1911).
S ar e B ue a eaf Kallima knyvettii de Ni
fl Hypolimnas bolina jacintha
rur
Recorded from NEN compound and Erule riverside
at Chizami. Uncommon. Reported to be common at low
elevations from the Naga Hills (Tytler 1911).
ea o
Cho o a e
ans Junonia iphita iphita Cramer
Recorded from many different locations at Chizami.
Seen in both pre- and post-monsoon seasons. Very
common. Reported to be very common from the Naga
Hills (Tytler 1911).
emon ans Junonia lemonias lemonias innaeus
Recorded from NEN compound between January
and July. Uncommon. Reported to be very common
from the low elevations of Naga Hills and not collected
above 1,200 m (Tytler 1911).
B ue ans Junonia orithya ocyale
ner
Recorded from many different locations at Chizami in
February, July and October. Very common. Reported to
be very common from the Naga Hills (Tytler 1911).
ndian B ue dmira Kaniska canace canace
innaeus
Recorded from many different locations at Chizami.
Common. Reported to be common from the Naga Hills
(Tytler 1911).
i e
Recorded from the Thetsumi trail at Chizami in July
2011. Rare. This species is legally protected under
Schedule II of the Wildlife (Protection) Act, 1972.
Reported to be “not uncommon” from the Naga Hills
(Elwes 1891). Tytler (1911) reports it to be rather rare
from the Naga Hills.
rea
2012, Chizami. Rare in the survey area. Reported to be
fairly common from the low elevations of Naga Hills
(Tytler 1911).
ans Junonia almana almana innaeus
Recorded from many different locations at Chizami
during the summer. Uncommon. Reported to be not
uncommon from the Naga Hills (Tytler 1911).
e o ans Junonia hierta hierta a ri ius
Recorded only one individual at Erule River in October
S otted es er Symbrenthia hypselis cotanda
oore
A single record from the NEN compound in August
2011. Rare. Reported to be rather common from the
Naga Hills (Tytler 1911).
Common es er Symbrenthia lilaea khasiana
oore
Recorded from the NEN compound, the Government
high school area and many other different parts of
Chizami through most of the year. Common. Reported
to be common from the Naga Hills (Tytler 1911).
B ue ai ed es er Symbrenthia niphanda niphanda
oore
A single record from the NEN compound in June
2013. Very rare. This species is legally protected
under Schedule II of the Wildlife (Protection) Act, 1972.
Reported to be rare in the Naga Hills with only three
individuals being collected in two years in August and
October (Tytler 1911).
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
ain ed ad Vanessa cardui cardui innaeus
Recorded from different locations of Chizami during
the summer season. Common. Reported to be very
common from the Naga Hills (Tytler 1911).
un e
or Thaumantis diores diores ou eda
A single record from Chizami Village late evening in
November 2012. Very rare in the survey area. Reported
from the Naga Hills (Tytler 1911).
ndian ed dmira Vanessa indica indica er s
Recorded from many different locations at Chizami
during the pre- and post- monsoon seasons. Common.
Reported to be common from the Naga Hills (Tytler
1911).
ami N m ha idae Su fami seuder o inae
Cons a e Dichorragia nesimachus nesimachus
o re
A single record from below the NEN compound in
August 2011. Very rare in the survey area.
a
Pseudergolis wedah wedah o ar
Recorded from Thetsumi and Erule trails at Chizami
mostly post- monsoon from July onwards. Often seen
mud-puddling. Common. Reported to be not uncommon
from the Naga Hills (Tytler 1911).
o in a Stibochiona nicea nicea ra
Records from the Thopfu and Tsetsumi trails at
Chizami in February 2010 and July 2011. Rare in the
survey area. Reported to be not uncommon in the Naga
Hills (Tytler 1911).
ami N m ha idae Su fami Sa rinae
e o
r ad Aemona amathusia e i son
Recorded from the NEN compound area at Chizami
in the summer. Uncommon. This species is legally
protected under Schedule II of the Wildlife (Protection)
Act, 1972. Reported to be very rare in the Naga Hills
(Tytler 1911).
Callerebia orixa oore
Recorded from the NEN compound. A few worn and
poorly marked individuals were recorded and could not
be identified to the species level with certainty, though
it is likely that these were C. orixa. Rare in the survey
area. Several specimens collected from the Naga Hills
and reported to be common post-monsoon (Elwes 1891;
Tytler 1911).
Common a mfl Elymnias hypermnestra undularis
rur
Recorded from Erule and Kuwari at Chizami during
the summer months. Common. Reported to be common
in the Naga Hills (Tytler 1911).
S otted a mfl Elymnias malelas malelas
e i son
Recorded from many different parts of Chizami’s
forests. Common.
Banded ree ro n Lethe confusa confusa
uri i ius
Recorded below the NEN compound in Mar 2013.
Rare. Reported to be common from Kohima (Tytler
1911).
Common ree ro n Lethe rohria rohria a ri ius
Recorded from many different locations at Chizami
between March and October. Common. Reported to be
common from Kohima (Tytler 1911).
ood
S rai h anded ree ro n Lethe verma sintica
ruhs orfer
Recorded from many different locations of Chizami
forests between March and October. Common. Seen
visiting animal sheds. Reported to be very common
at altitudes between 1,500–2,000 m in the Naga Hills
(Tytler 1911).
A single record from the Tahulu trail at Chizami in
September 2011. Very rare in the survey area. This
species is legally protected under Schedule II of the
Wildlife (Protection) Act, 1972.
n ed ed ores er Lethe chandica chandica
oore
Recorded from inside the village and from the NEN
compound in July 2011 and March 2013. Rare.
Common
u er Discophora sondaica zal
es ood
A single record from the NEN compound in May 2012.
Very rare in the survey area. Reported to be common
from low elevations in the Naga Hills (Tytler 1911).
ar e aun Faunis eumeus assama
es
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
Common ed ores er Lethe mekara mekara
oore
A single record from the NEN compound at Chizami
in October 2012. Very rare in the survey area.
ar
enin
Bro n Melanitis phedima bela
oore
Recorded from the NEN compound in July 2011 and
October 2012. Uncommon. Reported to be common in
the Naga Hills (Tytler 1911).
ai ed ed ores er Lethe sinorix sinorix e i son
Recorded from many different locations at Chizami
sporadically throughout the year including the winter
months. Common. This subspecies is legally protected
under Schedule II of the Wildlife (Protection) Act, 1972.
Reported from the Naga Hills (Elwes 1891; Tytler 1911).
Bam oo ores er Lethe f kansa oore
Two sightings of this butterfly in the same locationjust outside NEN compound , near a clump of bamboo
in January 2013. UNH ocellus in four on an even curve
hence seems to be Lethe kansa in its dry season form.
However, the upperside could not be seen, and hence
this identification is only provisional. Rare. Reported
from the Naga foothills from May to August (Tytler
1911).
a e ores er Lethe latiaris latiaris e i son
Recorded from the NEN compound at Chizami and
the forest trail below NEN compound in April 2011,
October 2012 and August 2013. Rare. Reported to be
not uncommon between 1,500–2,000 m, September
and October (Tytler 1911).
Common
ood ro n Lethe sidonis
rea
enin Bro n Melanitis zitenius zitenius
er s
A few records of individuals identified in hand in
DSF in forest below NEN compound in Oct 2012. Rare.
This species is legally protected under Schedule II of the
Wildlife (Protection) Act, 1972.
hi e ine Bush ro n Heteropsis malsara
oore
Recorded from many different locations of the
forested areas at Chizami and at the NEN compound in
July 2011 and October 2012. This species was earlier
considered to be part of the Genus Mycalesis but recent
phylogenetic studies have re-classified them in Genus
Heteropsis (Kodandaramaiah et al. 2010). Common.
Only a single record from the Naga Hills (Tytler 1911).
i a ine Bush ro n Mycalesis francisca sanatana
oore
Recorded from many different locations at Chizami in
July and October. Common. Seen mud-puddling at the
riverside occasionally, but seen mostly on the footpaths
leading to the fields and inside the bushes. Reported
from the Naga Hills (Elwes 1891, Tytler 1911).
e i son
Two individuals recorded from the NEN compound in
the summer months. Rare in the survey area. Reported
from the Naga Hills (Elwes 1891). Reported to be very
common at altitudes between 1,500–2,000 m in the
Naga Hills (Tytler 1911).
i a for Lethe f sura ou eda
A single unconfirmed record on the Thopfu trail
at Chizami in November 2012. The butterfly was
photographed sitting on the upper surface of a leaf,
high up in the canopy, and its upperside could not be
observed. Reported to be very common in the Naga Hills
(Tytler 1911).
Common
enin Bro n Melanitis leda leda
innaeus
Recorded from different locations at Chizami in the
pre- and post- monsoon seasons. Common. Reported to
be fairly common in the Naga Hills (Tytler 1911).
e ha Bush ro n Mycalesis lepcha kohimensis
er
Recorded from many different locations of the
forested areas at Chizami. Common. Dry season form
reported in March and April and wet season form in
August and October from Kohima (Tytler 1911; Tytler
1914).
hi e ed ed Bush ro n Mycalesis mestra sadona
er
Recorded from two locations at Chizami—the NEN
compound and at Awu in October 2012. Rare in the
survey area. This species is legally protected under
Schedule II of the Wildlife (Protection) Act, 1972.
eined a
rin h Neope pulaha pulaha
oore
Recorded from the NEN compound, near the
kitchen. Two individuals recorded at the same location
in September 2011 and May 2013. Rare. Reported from
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
i er Bro n Orinoma damaris damaris ra
Two individuals recorded from different locations at
Chizami in October 2012. Rare. Reported at 1,500m in
September and October and reported to be not common
in the Naga Hills (Tytler 1911).
ai ed a Graphium agamemnon agamemnon
innaeus
Recorded by Kewekhrozo Thopi on the village
approach road leading to Chizami Village near NEN area
and by SS in October 2012. Rare in the survey area.
Reported to be common from the Naga Hills (Tytler
1912).
Common i e rin Ypthima baldus baldus
a ri ius
Recorded from all locations at Chizami mostly
between March and July. Common. Reported to be
common from the Naga Hills (Tytler 1911).
ass B ue ott e Graphium cloanthus cloanthus
es ood
Recorded from the NEN compound. Often seen
visiting flowers during the summer. Rare. Reported to
be not common from the Naga Hills (Tytler 1912).
Ne ar hree rin Ypthima cf. newara newara
oore
Recorded from all locations at Chizami area in July
and October. Common. All the individuals recorded had
a larger sub-apical ocellus UNH and lacked UNF/H submarginal dark lines, than individuals recorded elsewhere
in northeastern India by SS. It is difficult to separate Y.
newara and Y. confusa without examining male genitalia
hence only provisionally identified.
Common B ue ott e Graphium sarpedon sarpedon
innaeus
Recorded from all locations at Chizami. Mostly seen
visiting flowers and mud-puddling on the riverbank.
Uncommon. Reported to be common from the Naga
Hills (Tytler 1912).
the Naga Hills (Elwes 1891).
ima a an i e rin Ypthima sakra sakra
oore
Recorded from all locations at Chizami in the preand post-monsoon seasons. Common. Reported from
the Naga Hills (Elwes 1891; Tytler 1911).
Ypthima s
At least two separate individuals, which did not match
any of the large-sized fiverings: Ypthima savara, Ypthima
methora, Ypthima dohertyi or Ypthima persimilis. Both
these individuals had wingspans > 45 mm, were heavily
striated on both wings UNF/H. UNF: in addition to large
sub-apical doubled-pupilled ocellus, and additional
ocellus in two. UNH: with five mostly equal-sized ocelli,
with pupils. Tornal ocelli merged. Seen at Chizami in the
forested areas in Oct 2012.
ami
a i ionidae Su fami a i ioninae
reen ra on ai Lamproptera meges indistincta
er
Recorded from all locations at Chizami. Mostly
seen during the summer between March and July.
Often seen mud-puddling. Common. Reported to be
not uncommon between 1,200–1,500 m from April to
October (Tytler 1912).
Common ime Papilio clytia clytia innaeus
Form dissimillis recorded from the Erule riverbank
at Chizami. Seen mud-puddling. Rare. Form dissimilis
reported to be common from the Naga Hills (Tytler
1912).
esser
ime Papilio epycides epycides e i son
A single record from the NEN compound in April
2014. Very rare.
ed e en Papilio helenus helenus innaeus
Recorded from all locations at Chizami between
March and October. Mostly seen visiting flowers. Locally
common. Reported to be common from the Naga Hills
(Tytler 1912).
e o
e en Papilio nephelus chaon
es
ood
Recorded from the Erule riverbank at Chizami in
August 2013. Rare. Reported at low elevations from the
Naga Hills (Tytler 1912).
rea
ormon Papilio memnon agenor innaeus
Recorded from the Erule riverbank and other
forested areas at Chizami in July and October. Seen mudpuddling, Forms agenor and distantianus seen. Rare.
Reported “sparingly” from the Naga Hills (Tytler 1912).
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Common
Naro & Sondhi
ormon Papilio polytes romulus Cramer
Recorded from the Awelu trail at Chizami. Rare in
the survey area. Reported to be common from the Naga
Hills (Tytler 1912).
ed reas Papilio alcmenor alcmenor e der &
e der
Recorded from the Erule riverbank at Chizami. Seen
mud-puddling. Rare. Reported from the Naga Hills
(Tytler 1912).
S an e Papilio protenor euprotenor ruhs orfer
(Tytler 1912).
Bird in s
Troides s
A birdwing seen in flight by SS though species level
identification was not possible.
ami
ieridae Su fami Co iadinae
hree s o rass e o Eurema blanda silhetana
a a e
Recorded from all locations at Chizami. Common.
Reported to be not common from August to September
at Kohima (Tytler 1912).
Recorded from all locations at Chizami. Seen mostly
in the months of March and July. Twenty-five individuals
were recorded in March alone. Very common. Reported
to be common from the Naga Hills (Tytler 1912).
Sma
rass e o Eurema brigitta rubella
a a e
Recorded from all locations at Chizami. Common.
Reported to be not very common from the Naga Hills
(Tytler 1912).
ime Butterfl Papilio demoleus demoleus
innaeus
Recorded from all locations at Chizami mostly during
the summer season. Common. Reported from the Naga
Hills (Tytler 1912).
Common rass e o Eurema hecabe hecabe
innaeus
Recorded from all locations at Chizami. Common.
Reported to be very common from the Naga Hills (Tytler
1912).
B ue ea o
Papilo arcturus arcturus
es
ood
Recorded from the NEN compound in October 2012.
Rare. Reported to be not uncommon from the Naga Hills
(Tytler 1912).
Common
ea o
Papilio bianor ganesa
ou eda
Recorded from numerous locations at Chizami.
Common. Reported rather commonly from the Naga
Hills (Tytler 1912).
aris ea o Papilio paris paris innaeus
Recorded from all locations at Chizami. Seen mostly
visiting flowers in gardens. Common. Reported to be
very common from the Naga Hills (Tytler 1912).
S o ess rass e o Eurema laeta sikkima
oore
Recorded from many different locations at Chizami.
Common. Reported to be rather common from Kohima
(Tytler 1912).
ami
ieridae Su fami ierinae
Common
a ross Appias albina darada e der &
e der
Recorded from the NEN compound. Male and female
seen. Rare in the survey area. This subspecies is legally
protected under Schedule II of the Wildlife (Protection)
Act, 1972. Reported to be rare from the Naga Hills
(Tytler 1912).
Common Ba in Atrophaneura varuna astorion
es ood
Recorded from the NEN compound. Rare. Collected
“sparingly” in the Naga Hills (Tytler 1912).
ran e
a ross Appias galba a a e
Two individuals recorded from Chizami Village. Seen
mud-puddling. Rare. This subspecies is legally protected
under Schedule IV of the Wildlife (Protection) Act, 1972.
Reported to be not uncommon from the Naga Hills
(Tytler 1912).
Common indmi Byasa polyeuctes polyeuctes
ou eda
Recorded from many different locations at Chizami.
Common. Collected “commonly” from the Naga Hills
S o u n Appias lalage lalage ou eda
Recorded from many different locations at Chizami.
Common. Reported to be rather common from the
Naga Hills (Tytler 1912).
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
esser u Cepora nadina nadina u as
Recorded from two locations—the NEN compound
and Erule riverbank at Chizami. Rare in the survey area.
A few individuals recorded from the Naga Hills (Tytler
1912).
Common mi ran Catopsilia pomona pomona
a ri ius
Recorded from the NEN compound. Uncommon.
Reported to be not uncommon from the Naga Hills
(Tytler 1912).
a e anderer Pareronia avatar oore
Recorded from different locations of Chizami.
Common. A few individuals reported from the Naga Hills
(Tytler 1912).
ar e Ca a e hi e Pieris brassicae nepalensis
ou eda
Recorded from all locations at Chizami through most
of the year. Very common. Reported to be very scarce
from the Naga Hills (Tytler 1912).
ndian Ca
ott ed mi ran Catopsilia pyranthe pyranthe
innaeus
Recorded from the NEN compound. Uncommon.
ed reas eze e Delias acalis pyramus
i eze e Delias belladonna lugens ordan
Recorded from many different parts at Chizami.
Common. Recorded as common between 1,200–2,100
m in August and September in the Naga Hills (Tytler
1912).
ed s o eze e Delias descombesi descombesi
Boisdu a
A single record from Chizami Village in November
2012. Very rare. A few individuals reported from the
Naga Hills (Tytler 1912).
ed ase eze e Delias pasithoe pasithoe
innaeus
Recorded from the NEN compound and the Erule
riverbank at Chizami during summer. Rare.
rea ran e
Hebomoia glaucippe glaucippe
innaeus
Recorded from many different locations of Chizami.
Common. Reported to be common from low elevations
up to 600m in the Naga Hills (Tytler 1912).
ran e
hi e Pieris canidia indica
ans
Recorded from all locations at Chizami through most
of the year. Very common. Reported to be abundant
throughout the year from the Naga Hills (Tytler 1912).
a a e
Recorded from the NEN compound and the Erule
riverbank at Chizami in October 2012. Rare.
e o
a e
Ixias pyrene familiaris Bu er
Recorded from many different locations at Chizami
between March and October. Common. Reported to be
common from the Naga Hills (Tytler 1912).
Ba h hi e Pontia daplidice moorei
er
Recorded from many different parts of Chizami.
Common. This species has extended its range eastwards
in recent times from the western and central Himalaya
to many locations in eastern Himalaya including Assam,
Arunachal Pradesh, Nagaland and Manipur (Naro &
Sondhi 2013; Singh & Gogoi 2013).
reen eined
hi e Pieris melete ajaka
oore
Recorded from all locations at Chizami through
most of the year. Very common. Reported to be rather
uncommon from the Naga Hills (Tytler 1912).
S otted Sa oo h Prioneris thestylis thestylis
ou eda
Recorded from many different parts at Chizami.
Female recorded feeding on flower nectar in July 2011
in the NEN office complex. Males common, females
rare. Reported to be common from the Naga Hills (Tytler
1912).
ami
iodinidae Su fami Nemeo iinae
ar ud Abisara fylla es ood
Recorded from all locations at Chizami. Seen
throughout the year. Very common. Reported to be
abundant and common from the Naga Hills (Elwes 1891;
Tytler 1911).
S ri ed un h Dodona adonira adonira e i son
A single record from Thetsumi roadside near the
stream in February 2013. Rare. Reported to be “not
common” from the Naga Hills (Elwes 1891; Tytler 1911).
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
un hine o Zemeros flegyas fleygas Cramer
Recorded from all locations at Chizami. Seen
throughout the year. Very common. Reported to be
very common from the Naga Hills (Tytler 1911).
NC S
Bin ham C
The Fauna of British India including Ceylone
and Burma: Butterflies. Vols. I-II. Taylor and Francis Ltd., London.
Cor e
S &
end e ur
The Butterflies of the Malay
Peninsula. 4th Edition revised by J.N. Eliot. Malayan Nature Society,
Kuala Lumpur, 595pp.
de on
Revision of the oriental Genus Matapa Moore
(Lepidoptera: Hesperiidae) with discussion of its phylogeny and
geographic history. Zoologische Mededelingen 57: 21.
de Ni i e C
On new and little-known Lepidoptera from
the Oriental Region. Journal of Bombay Natural History Society
13(1): 157–175.
io
N
An Analysis of the Eurasian and Australian Neptini
(Lepidoptera: Nymphalidae). Bulletin of the British Museum
(Natural History) Entomology. Supplement 15.
es
On butterflies collected by Mr. W. Doherty in the
Naga and Karen Hills and in Perak. Proceedings of the Zoological
Society of London 1891: 249–289.
es
On butterflies collected by Mr. W. Doherty in the
Naga and Karen Hills and in Perak. Proceedings of the Zoological
Society of London 1892: 617–664.
ans
The Identification of Indian Butterflies. 2nd Revised
Edition. Bombay Natural History Society, Bombay, x+454pp+32pl.
ans
A catalogue of the Hesperiidae from Europe,
Asia and Australia in the British Museum (Natural History). British
Museum (Natural History), London, 502pp.
ans
A revision of the Arhopala group of Oriental
Lycaenidae (Lepidoptera: Rhopalocera). Bulletin of the British
Museum (Natural History), London Entomology Volume 5: 85–141.
ari a
The Butterflies of The Sikkim Himalaya and Their
Natural History. Sikkim Nature Conservation Foundation, Gangtok,
217pp.
uan
&
ue
The Chinese Pseudocoladenia skippers.
Neue Entomologische Nachrichten 57: 161–170.
ehim ar
The Book of the Indian Butterflies. Bombay Natural
History Society and Oxford University Press, Oxford, xvi+497pp.
in on S
Illustrated Checklist for the Butterflies of Myanmar.
Smithsonian Institution, 197pp.
odandaramaiah
C ees C
er
orres
aran h
& N
ah er
Phylogenetics and biogeography of a
spectacular Old World radiation of butterflies: the subtribe
Mycalesina (Lepidoptera: Nymphalidae: Satyrini). BMC Evolutionary
Biology 10: 172.
un e
Butterflies of Peninsular India. Universities Press
(Hyderabad) and Indian Academy of Sciences (Bangalore), 254pp.
arsha
&
de Ni i e
The Butterflies of India,
Burmah and Ceylon. Vol. I. The Calcutta Central Press Co., Calcutta,
398pp
arsha
&
de Ni i e
The Butterflies of India,
Burmah and Ceylon. Vol. II.The Calcutta Central Press Co., Calcutta,
370pp.
arsha
&
de Ni i e
The Butterflies of India,
Burmah and Ceylon. Vol. III. The Calcutta Central Press Co., Calcutta,
552pp
Naro & Sondhi
Naro
Sighting of Aglais cashmirensis aesis Fruhstorfer, 1912
(Nymphalidae) from Nagaland, India. Journal of Threatened Taxa
4(4): 2534–2535; http://dx.doi.org/10.11609/JoTT.o3019.2534-5
Naro
& S Sondhi
Sightings of Bath White Pontia daplidice
moorei Röber, 1907 (Lepidoptera: Pieridae: Pierinae: Pierini) from
Arunachal Pradesh and Nagaland, India. Journal of Threatened Taxa
5(7): 4122-4124; http://dx.doi.org/10.11609/JoTT.o3308.4122-4
Sin h
Butterflies of India. Om Books International, 183pp.
Sin h
&
o oi
A new range record for the Bath White
butterfly Pontia daplidice for north-east India. Bionotes 15(2): 59.
Smi h C
Illustrated Checklist of Nepal’s Butterflies. T.C.
Majupuria (Ed.). Rohit Kumar, Gwalior. M.P., India, 127pp.
Smi h C
Butterflies of Nepal. Revised Edition, Tecpress Service
L.P., Bangkok, Thailand, 368pp.
Sondhi S &
un e
Butterflies and Moths of Pakke Tiger
Reserve. Titli Trust (Dehradun) and Indian Foundation for Butterflies
(Bengaluru), vi+202pp.
Sondhi S
un e
a e ar
o a e ar &
o e ar
Butterflies of the Garo Hills. Samrakshan Trust (New Delhi), Titli
Trust (Dehradun), and Indian Foundation for Butterflies (Bengaluru),
xvi+200pp.
S inhoe C
Lepidoptera Indica. Vol. X. Rhopalocera.
Family Hesperiidae. Reeve & Co, London, 361pp+78pl.
a o
The Fauna of British India, including Ceylone
and Burma: Butterflies. Vol. 1. Taylor and Francis, London,
600pp+3pl.
a o
The Fauna of British India, including Ceylone and
Burma: Butterflies. Vol. 2. Taylor and Francis, London, 506pp+2pl.
er C
Notes on butterflies from the Naga Hills - Part II.
Journal of Bombay Natural History Society 21: 588–606.
er C
Notes on butterflies from the Naga Hills - Part I.
Journal of Bombay Natural History Society 21: 48–65.
er
C
Notes on some new and interesting butterflies
from Manipur and the Naga Hills. Part - I. Journal of Bombay Natural
History Society 23: 216–229.
er C
a Notes on some new and interesting butterflies
from Manipur and the Naga Hills. Part - II. Journal of Bombay
Natural History Society 23: 502–515.
er C
Notes on some new and interesting butterflies
from Manipur and the Naga Hills. Part - III. Journal of Bombay
Natural History Society 24: 119–155.
er
C
Notes on Some New and Interesting Butterflies
chiefly from Burma. Part - 2. Journal of Bombay Natural History
Society 42: 109–123.
arshne
a First Bibliography of the Butterflies of Indian
Region. Bionotes 10(2): 43–46.
arshne
First Bibliography of the Butterflies of Indian
Region. Bionotes 10(3): 81–86.
arshne
First Bibliography of the Butterflies of Indian
Region. Bionotes 10(4): 143–146.
arshne
a First Bibliography of the Butterflies of Indian
Region. Bionotes 11(1): 30–33.
arshne
First Bibliography of the Butterflies of Indian
Region. Bionotes 11(2): 47–50.
arshne
First Bibliography of the Butterflies of Indian
Region. Bionotes 11(3): 86–89.
arshne
d First Bibliography of the Butterflies of Indian
Region. Bionotes 11(4): 119–121.
n er B h
Butterflies of The Indian Region. Bombay
Natural History Society, Bombay, xx+523pp+72pl.
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
ma e
Sae
ma e ndian
japonica
Badamia exclamationis
reen
e Bibasis vasutana
(©Tshetsholo Naro NEN)
ma e
in Choaspes benjaminii
(©Tshetsholo Naro NEN)
Bro n
(©Tshetsholo Naro NEN)
ma e
(© Sanjay Sondhi)
(© Kewekhrozo Thopi NEN)
(©Tshetsholo Naro NEN)
Hasora anura anura
(©Tshetsholo Naro NEN)
ma e
o
er
Hasora tamina-
er Astictopterus jama
S rai h S i Parnara f guttatus
(©Tshetsholo Naro NEN)
ma e
ma e
ores
olivascens
o
ur e S i Caltoris tulsi
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
eined S ru o er
Aeromachus stigmata obsoleta
ma e
hi e anded
tus bhavara
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e B ue s otted S ru
Aeromachus kali
ma e
sinensis
ar e Banded S i Pelopidas
ma e
Sma Banded S i Pelopidas
mathias mathias
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
e o S o S i Polytremis eltola
ma e
ar s
Potanthus s
(©Tshetsholo Naro NEN)
ma e
eltola
(©Tshetsholo Naro NEN)
(© Sanjay Sondhi)
ma e
ssam ar er Ochlodes siva siva
ma e
u ed
e Sebastonyma dolopia
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ssam ar er Ochlodes siva siva
ma e
i h S ra
e Pithauria
stramineipennis stramineipennis
(©Tshetsholo Naro NEN)
ma e
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
© Kewekhrozo Thopi NEN
ma e
Ba
eined ed e e
Matapa f sasivarna
ma e
(© Sanjay Sondhi)
ndian a m Bo Suastus gremius
(© Kewekhrozo Thopi NEN)
(©Tshetsholo Naro NEN)
ma e
gremius
Si im a m ede e Erionota torus
ma e
es ri ed emon Notocrypta
curvifascia curvifascia
ma e
S otted emon Notocrypta
feisthamelii alysos
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
ma e
Common Banded emon
Notocrypta paralysos asawa
Butterflies of Chizami
Naro & Sondhi
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
Common S otted a
Celaenorrhinus leucocera
ma e
udd ied a
Pseudocoladenia f fatua
ree itter Hyarotis adrastus
(©Tshetsholo Naro NEN)
ma e
praba
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
agama
S otted n e Caprona agama
ma e
Bri h Sun eam Curetis bulis bulis
ma e
vasava
a Tagiades
(©Tshetsholo Naro NEN)
ma e
S otted Sno
menaka menaka
n e Ctenoptilum vasava
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
a n
(© Kewekhrozo Thopi NEN)
ma e
o den Sa
brahma brahma
ma e
Bri h Sun eam Curetis bulis bulis
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
n e Odontoptilum
(©Tshetsholo Naro NEN)
ma e
Ches nu
angulata angulata
ma e
Na a Sa hire Heliophorus
kohimensis kohimensis
hire Heliophorus
ma e
Na a Sa hire Heliophorus
kohimensis kohimensis
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Naro & Sondhi
ma e
reen Sa
moorei tytleri
hire Heliophorus
ma e
oin ed Ci ia e B ue Anthene
lycaenina lycambes
(©Tshetsholo Naro NEN)
ma e
Na a Sa hire fema e
Heliophorus kohimensis kohimensis
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ma e
ain ed e B ue Celastrina
lavendularis limbata
(© Sanjay Sondhi)
ma e
a e ed e B ue Udara dilecta
ma e
ed e Cu id Bothrinia chennellii
Bothrinia chennellii ed e Cu id
ed e Cu id Bothrinia chennellii
(©Tshetsholo Naro NEN)
ma e
ma e
dilecta
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
Common ed e B ue Acytolepis
puspa gisca
ma e
Cha man s Cu id Everes argiades
diporides
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Image 42. Silver Forget-me-not
Catochrysops panormus exiguus
Image 41. Indian Cupid Everes lacturnus
assamica
(©Tshetsholo Naro NEN)
Image 41. Indian Cupid Everes lacturnus
assamica
(© Kewekhrozo Thopi NEN)
Naro & Sondhi
(© Kewekhrozo Thopi NEN)
Butterflies of Chizami
Image 44. Gram Blue female Euchrysops
cnejus cnejus
(©Tshetsholo Naro NEN)
Image 45. Dark Cerulean Jamides bochus
bochus
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Image 43. Forget-me-not
Catochrysops strabo strabo
Image 46. Cerulean Jamides sp.
Image 47. Glistening Cerulean Jamides elpis
pseudoelpis
Image 48. Pea Blue Lampides boeticus
6616
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
© Kewekhrozo Thopi NEN
in
ine ue Petrelaea dana
(©Tshetsholo Naro NEN)
ar
rass B ue Zizeeria karsandra
o a Ancema ctesia ctesia
ma e
watsoni
ai ess Bush ue Arhopala ganesa
ma e
eaf B ue Amblypodia anita dina
UN
UP
(©Tshetsholo Naro NEN)
Bi S o
(©Tshetsholo Naro NEN)
ma e
Common ine ue Prosotas nora
ma e
Common em Poritia hewitsoni
hewitsoni
(©Tshetsholo Naro NEN)
ma e
ma e
ardates
(©Tshetsholo Naro NEN)
ma e
UN
(©Tshetsholo Naro NEN)
e ra B ue Leptotes plinius plinius
(©Tshetsholo Naro NEN)
ma e
(©Tshetsholo Naro NEN)
Naro & Sondhi
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ma e
ma e
o dered a
bazalus teetsa
ue Arhopala
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
UP
Comi
a
ue Arhopala comica
Naro & Sondhi
UN
ue
ma e
Common inse Catapaecilma
major major
(©Tshetsholo Naro NEN)
a
UP
© Kewekhrozo Thopi NEN
ai ess
ush ue Flos areste
(©Tshetsholo Naro NEN)
ma e
ma e
Si era rea B ue Iraota timoleon
timoleon
© Kewekhrozo Thopi NEN
ma e
Common ash Rapala nissa
nissoides
(©Tshetsholo Naro NEN)
ma e
e o dis
Arhopala singla
(©Tshetsholo Naro NEN)
(© Khromese-u Thopi)
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ma e
ulupi
ma e
ambica
ur e m eror Apatura ambica
a n
ma e
Common a ia B ue
Surendra quercetorum quercetorum
(©Tshetsholo Naro NEN)
ha i Si er ine Spindasis rukmini
(©Tshetsholo Naro NEN)
ma e
(©Tshetsholo Naro NEN)
ma e
C u Si er ine Spindasis syama
peguanus
m eror Chitoria ulupi
ma e
ulupi
a n
m eror Chitoria ulupi
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Naro & Sondhi
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ma e
(©Tshetsholo Naro NEN)
o den m eror Dilipia morgiana
ma e
Ser ean m eror
Mimathyma chevana chevana
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
Ser ean m eror
Mimathyma chevana chevana
ma e
Cour esan Euripus nyctelius
nyctelius
ma e
ma e
asha Herona marathus marathus
Cir e Hestinalis nama nama
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
chandra
Siren Hestina persimilis persimilis
rin e Rohana parisatis
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
Ba
parisatis
as ern Cour er Sephisa chandra
ma e
Common Cas or Ariadne merione
tapestrina
(©Tshetsholo Naro NEN)
ma e
ma e
rea Na a Charaxes
eudamippus eudamippus
ma e
Common Na a Charaxes
athamas athamas
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
Common
thyodamas
(©Tshetsholo Naro NEN)
ma e
ain i er Danaus chrysippus
chrysippus
ma e
genutia
S ri ed i er Danaus genutia
(©Tshetsholo Naro NEN)
a Cyrestis thyodamas
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
S ri ed B ue Cro fema e
Euploea mulciber mulciber
ma e
ass
melanoides
(©Tshetsholo Naro NEN)
ma e
Parantica sita sita Ches nu
ma e
ar B ue i er Tirumala
septentrionis septentrionis
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
Cho o a e i er Parantica
melaneus plataniston
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
S ri ed B ue Cro Euploea mulciber mulciber
i er Parantica aglea
i er
© Kewekhrozo Thopi NEN
ma e
issoria
e o Cos er Acraea issoria
ma e
childreni
ar e Si ers ri Argynnis childreni
ma e
ro i a ri
hyperbius hyperbius
ar Argynnis
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Naro & Sondhi
(©Tshetsholo Naro NEN)
ar e eoman Cirrochroa aoris
(©Tshetsholo Naro NEN)
ma e
ed a e in fema e
Cethosia biblis tisamena
ma e
ed a e in ma e
Cethosia biblis tisamena
ma e
aoris
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ma e
mithila
female
male
ma e
cyane
eo ard a e in Cethosia cyane
Common eoman Cirrochroa tyche
male
UP
ma e
Common eo ard
Phalanta phalantha phalantha
(©Tshetsholo Naro NEN)
ma e
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
UN
us
female
ma e
Cupha erymanthis lotis
ma e
a ran Vagrans egista sinha
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Cruiser Vindula erota erota
Naro & Sondhi
Common Bea Libythea lepita
(©Tshetsholo Naro NEN)
rea Calinaga brahma brahma
ren h u e Euthalia franciae
(©Tshetsholo Naro NEN)
ma e
S a Ser ean Athyma selenophora selenophora
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
ran e S a Ser ean fema e
Athyma cama cama
ma e
B ue u e Bassarona durga
splendens
(©Tshetsholo Naro NEN)
ma e
B ue u e Bassarona durga
splendens
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
franciae
Common Ser ean Athyma perius
(©Tshetsholo Naro NEN)
ma e
ran e S a Ser ean ma e
Athyma cama cama
ma e
ma e
perius
ma e
ima a an Ser ean Athyma
opalina orientalis
(©Tshetsholo Naro NEN)
ma e
lepita
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ma e
Bronze u e Euthalia nara
nagaensis
ma e
nadaka
reen u e Euthalia sahadeva
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
(© Sanjay Sondhi)
ma e
Common Baron Euthalia
aconthea garuda
(©Tshetsholo Naro NEN)
ma e
hi e ed ed B ue Baron
Euthalia phemius phemius
e o Sai or Neptis ananta
ma e
astola
Nam a Sai er Neptis f namba
(©Tshetsholo Naro NEN)
Nam a Sai er Neptis f namba
ma e
Broad anded Sai er Neptis
sankara amba
a as Sai er Neptis sappho
ma e
Broad anded Sai er Neptis
sankara amba
ma e
astola
a as Sai er Neptis sappho
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
namba
ma e
namba
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
ochracea
aud Baron Euthalia lubentina
aud Baron Euthalia lubentina
(©Tshetsholo Naro NEN)
ma e
lubentina
ma e
lubentina
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
procris
Commander Moduza procris
(©Tshetsholo Naro NEN)
ma e
Ci
gambrisius
(©Tshetsholo Naro NEN)
Naro & Sondhi
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ma e
ndian or oiseshe Aglais
caschmirensis aesis
er Parthenos sylvia
ma e
dohertyi
on o Araschnia prorsoides
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
u umn eaf Doleschallia
bisaltide indica
rea
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
jacintha
ma e
S ar e B ue a eaf
Kallima knyvettii
ma e
S ar e B ue a eaf
Kallima knyvettii
(©Tshetsholo Naro NEN)
ran e a eaf Kallima inachus
ran e a eaf Kallima inachus
(©Tshetsholo Naro NEN)
ma e
inachus
ma e
inachus
(©Tshetsholo Naro NEN)
on o Araschnia prorsoides
(©Tshetsholo Naro NEN)
ma e
dohertyi
fl Hypolimnas bolina
ma e
jacintha
rea
fl Hypolimnas bolina
ma e
almana
ea o
ans Junonia almana
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Naro & Sondhi
ans Junonia hierta
(©Tshetsholo Naro NEN)
ma e
iphita
Cho o a e ans Junonia iphita
ma e
ocyale
ma e
canace
B ue dmira Kaniska canace
ma e
lemonias
eoman ans Junonia lemonias
(©Tshetsholo Naro NEN)
e o
(©Tshetsholo Naro NEN)
ma e
hierta
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Butterflies of Chizami
B ue ans Junonia orithya
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
Common es er Symbrenthia
lilaea khasiana
(©Tshetsholo Naro NEN)
ma e
S otted es er Symbrenthia
hypselis cotanda
ma e
B ue ai ed es er Symbrenthia
niphanda niphanda
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
S otted es er Symbrenthia
hypselis cotanda
ma e
cardui
ma e
B ue ai ed es er Symbrenthia
niphanda niphanda
(©Tshetsholo Naro NEN)
ain ed ad Vanessa cardui
ma e
ndian ed dmira
Vanessa indica indica
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
(©Tshetsholo Naro NEN)
ma e
Cons a e Dichorragia nesimachus nesimachus
a
Pseudergolis wedah
ma e
(©Tshetsholo Naro NEN)
ar e aun Faunis eumeus
or Thaumantis diores
(©Tshetsholo Naro NEN)
ma e
diores
un e
or Thaumantis diores
ma e
(©Tshetsholo Naro NEN)
un e
(©Tshetsholo Naro NEN)
ma e
diores
ma e
assama
r ad Aemona amathusia
(©Tshetsholo Naro NEN)
e o
o in a Stibochiona nicea nicea
© Kewekhrozo Thopi NEN
ma e
Common u er Discophora
sondaica zal
ma e
ma e
(©Tshetsholo Naro NEN)
ma e
wedah
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
© Kewekhrozo Thopi NEN
ma e
Common a mfl
Elymnias hypermnestra undularis
Callerebia orixa
(©Tshetsholo Naro NEN)
ma e
S otted a mfl
Elymnias malelas malelas
Callerebia ori a
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Butterflies of Chizami
Naro & Sondhi
(©Tshetsholo Naro NEN)
Image 152. Bamboo Forester Lethe cf. kansa
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Image 149. Angled Red Forester
Lethe chandica chandica
Image 150. Common Red Forester
Lethe mekara mekara
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Image 150. Common Red Forester
Lethe mekara mekara
Image 147. Common Treebrown
Lethe rohria rohria
Image 148. Straight-Banded Treebrown
Lethe verma sintica
(©Tshetsholo Naro NEN)
Image 148. Straight-banded Treebrown
Lethe verma sintica
Image 146. Banded Treebrown
Lethe confusa confusa
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Image 145. Spotted Palmfly
Elymnias malelas malelas
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Image 151. Tailed Red Forester
Lethe sinorix sinorix
(©Tshetsholo Naro NEN)
Image 153. Pale Forester Lethe latiaris
latiaris
Image 153. Pale Forester Lethe latiaris
latiaris
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
6627
ma e
Common
Melanitis leda leda
enin Bro n
ma e
Common
Melanitis leda leda
enin Bro n
(©Tshetsholo Naro NEN)
ood ro n
(©Tshetsholo Naro NEN)
ma e
Common
Lethe sidonis
i afor Lethe f sura
(©Tshetsholo Naro NEN)
ma e
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Naro & Sondhi
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ma e
ar
enin Bro n
Melanitis phedima bela
(©Tshetsholo Naro NEN)
ma e
ar
enin Bro n
Melanitis phedima bela
(© Sanjay Sondhi)
(© Sanjay Sondhi)
ma e
rea enin Bro n
Melanitis zitenius zitenius
ma e
hi e ine Brush ro n
Heteropsis malsara
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
rea enin Bro n
Melanitis zitenius zitenius
(©Tshetsholo Naro NEN)
ma e
sis malsara
hi e ine Brush ro n Heteropma e
i a ine Bush ro n
Mycalesis francisca sanatana
ma e
e ha Brush ro n
Mycalesis lepcha kohimensis
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Naro & Sondhi
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Butterflies of Chizami
(©Tshetsholo Naro NEN)
ma e
pulaha
ma e
hi e ed ed Bus
Mycalesis mestra sadona
rin h Neope pulaha
ro n
(© Sanjay Sondhi)
eined a
rin h Neope pulaha
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
pulaha
ma e
damaris
eined a
i er Bro n Orinoma damaris
ma e
Ne ar hree rin
Ypthima f newara newara
(© Sanjay Sondhi)
ma e
Common i e rin Ypthima
baldus baldus
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
ima a an i e rin Ypthima
sakra sakra
ma e
ima a an i e rin Ypthima
sakra sakra
© Kewekhrozo Thopi NEN
Ypthima s
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
ma e
reen ra on ai Lamproptera
meges indistincta
ma e
ai ed a Graphium agamemnon
agamemnon
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
ma e
ass B ue ott e Graphium
cloanthus cloanthus
Naro & Sondhi
Common
ime Papilio clytia
(©Tshetsholo Naro NEN)
esser
ime Papilio epycides
ma e
helenus
ed e en Papilio helenus
ma e
chaon
e o
e en Papilio nephelus
ime Papilio epycides
ma e
romulus
Common
ormon Papilio polytes
ma e
alcmenor
ed reas Papilio alcmenor
(©Tshetsholo Naro NEN)
ma e
demoleus
ma e
S an e Papilio protenor
euprotenor
(©Tshetsholo Naro NEN)
ormon Papilio memnon
(©Tshetsholo Naro NEN)
rea
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
agenor
esser
(©Tshetsholo Naro NEN)
ma e
Common B ue ott e Graphium
sarpedon sarpedon
ma e
epycides
ma e
epycides
(©Tshetsholo Naro NEN)
ma e
clytia
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ime Butterfl Papilio demoleus
ma e
arcturus
B ue ea o
Papilo arcturus
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
Naro & Sondhi
ma e
ma e
Common Ba
varuna astorion
Papilio bianor
(©Tshetsholo Naro NEN)
ma e
hree s o rass e o
Eurema blanda silhetana
ma e
Sma
brigitta rubella
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
Common indmi
Byasa polyeuctes polyeuctes
ma e
S o ess rass e o
Eurema laeta sikkima
ma e
Common rass e o
Eurema hecabe hecabe
in Atrophaneura
(©Tshetsholo Naro NEN)
Common ea o
Papilio paris paris
(©Tshetsholo Naro NEN)
ma e
ganesa
aris ea o
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
rass e o Eurema
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ma e
darada
Common
a ross Appias albina
ma e
ran e
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
a ross Appias galba
ma e
S o
u n Appias lalage lalage
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
ma e
Cho o a e
lyncida eleonora
a ross Appias
Naro & Sondhi
ma e
pyramus
ed reas eze e Delias acalis
(©Tshetsholo Naro NEN)
ma e
Common mi ran Catopsilia
pomona pomona
ma e
ott ed mi ran Catopsilia
pyranthe pyranthe
ma e
lugens
i
(©Tshetsholo Naro NEN)
esser u Cepora nadina nadina
(©Tshetsholo Naro NEN)
ma e
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
Butterflies of Chizami
eze e Delias belladonna
Hebomoia
ma e
familiaris
ma e
e o
ran e
Ixias pyrene
ae
anderer Pareronia avatar
ma e
ar e Ca a e
brassicae nepalensis
hi e Pieris
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ed ase eze e Delias pasithoe
(©Tshetsholo Naro NEN)
ma e
pasithoe
ma e
rea ran e
glaucippe glaucippe
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
(©Tshetsholo Naro NEN)
ma e
ed s o eze e Delias
descombesi descombesi
ma e
ndian Ca
Pieris canidia indica
a e
hi e
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
ar
ud Abisara fylla
reen iened
hi e Pieris melete
(©Tshetsholo Naro NEN)
ma e
S otted Sa
thestylis thestylis
oo h Prioneris
ma e
adonira
S ri ed un h Dodona adonira
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
(©Tshetsholo Naro NEN)
ma e
hi e Pontia daplidice
ma e
ajaka
(©Tshetsholo Naro NEN)
Ba h
(©Tshetsholo Naro NEN)
ma e
moorei
(©Tshetsholo Naro NEN)
Naro & Sondhi
(©Tshetsholo Naro NEN)
Butterflies of Chizami
ma e
fleygas
un hine o Zemeros flegyas
6633
Butterflies of Chizami
a
Naro & Sondhi
b
c
d
a
endi
rimari isi ed areas and heir ha i a s
she sho o Naro N N
e u rai
ru e i er an
ie of he sumi rai durin monsoon d
ie of Nor h as Ne
or
N N om ound
hrea ened a a
Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634
JOTT RECOGNIZES THIS CONSERVATION INITIATIVE
EMPOWERING YOUNGSTERS IN CONSERVING BIODIVERSITY - CHIZAMI
EXPERIENCE
The Hoolock Gibbon Eco Club (HGEC) was formed
in 2010. This was the first eco-club to be formed in
Nagaland and has been a pioneering effort of an
NGO called the North East Network (NEN). The
club has 50 children in the age group of 10–14 years
from the village of Chizami and the neighbouring
village of Enhulumi. HGEC was an outcome of a
wildlife education program to aid in mitigating
wild animal hunting titled “Under the Canopy” that
Dusty foot Productions (www.dustyfootindia.com)
and Go Wild (payalmolur.blogspot.in) conducted
in 2009. While filming a story on orphaned bear
cubs, the Dusty Foot Production team came across
rampant hunting and selling of wild meat in
northeastern India. This led them to team up with
a wildlife education group Go Wild Workshops to
create an education program and curriculum to shift
the focus from hunting to research, photography and
enquiry. North East Network and its tryst with
wildlife education and conservation began in 2009
when approached by the Rita Banerji (Dusty Foot
Productions) and Payal Molur (Go Wild).
North East Network (www.northeastnetwork.
org) is a woman’s rights organization in northeast
India, established in 1995. NEN’s role has been
that of a facilitator to empower women of northeast
around issues of livelihood, health, conflict and
governance through capacity building, awareness
raising, networking, research and advocacy. North
East Network and its tryst with wildlife education
and conservation began in 2009 when approached
by the Rita Banerji (Dusty Foot Productions) and
Payal Molur (Go Wild) to be the local partner. Titli
trust (www.titlitrust.com), was also brought in to
bring a new dimension to the scientific education
program. From there they have expanded the program
exponentially and now have trainers and clubs in
three districts of Nagaland.
The teachers and children trained in Nagaland,
have not only understood the importance of
conserving their forests but also have taken it to the
next stage of maintaining biodiversity registers
and cataloguing the animals that are around them
in a scientific manner.
Over the past four years the club has been
churning out a series of wildlife publications in the
form of posters, booklets, journal articles and popular
articles:
A few of the publications of HGEC
- First and second edition booklets with 171 and
217 species of butterflies of Chizami respectively
(2012 and 2014)
- A booklet on the Birds of Chizami (2014)
- Posters on wildlife of Chizami, Birds of Chizami
and Butterflies and Moths of Chizami
- Naro, T. (2012). Sighting of Aglais
cashmirensis aesis Fruhstorfer, 1912 (Nymphalidae)
from Nagaland, India. Journal of Threatened Taxa
4(4): 2534–2535; http://dx.doi.org/10.11609/
JoTT.o3019.2534-5
- Naro, T. & S. Sondhi (2013). Sightings of
Bath White Pontia daplidice moorei Röber, 1907
(Lepidoptera: Pieridae: Pierinae: Pierini) from
Arunachal Pradesh and Nagaland, India. Journal of
Threatened Taxa 5(7): 4122–4124; http://dx.doi.
org/10.11609/JoTT.o3308.4122-4
Payal Molur
payal.bmolur@gmail.com
Rita Banerji explaing the importance of lighting.
Chizami Village, Phek, Nagaland
Sanjay Sondhi describing the beauty of butterflies at the NEN
campus.
Understanding macro photography techniques
Photographing some animals is easier with a tripod
Diligently cataloging moths
Photo credits: Payal B Molur and Tshetsholo Naro
Payal Molur explaining how to map an area
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